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once and forms a short suspensor, so that all the organs of the 
young embryo, including the foot, are really of epibasal origin.” 
In a later publication (10) Campbell established the presence 
of the suspensor in the embryo of Macroglossum alidae. 
3. The late development and the position of the second root 
resemble closely the facts as they are in Danaea. Lyon remarks : 
“A study of the mature sporophyte of the ternatum species of 
Botrychium discloses unique characters which mark them as a 
natural group entitled to generic rank.” This opinion seems to 
be entirely justified by the facts. 
IV. 
Another representative of the ‘fernatum group, B. silaei- 
folium, was only available in the adult stage. This specimen 
was collected near Lake Tahoe, California, by Dr. Campbell 
and it possessed three full grown leaves 15—25 cm long and 
a fertile spike 730 cm long. The elaborately developed root 
system was also well preserved (Fig. LE Series of sections, 
both with hand microtome and with the paraffin method were 
made from petiole, bud and roots. 
The bundle in the petiole is very different from the 
other bundles which we found in other species of Botrychium. 
Its structure 1s essentially the same as that of the petiolar 
bundle in the fossil Osmundaceae (Bertrand, 2) although it 
shows in some points a marked relation to the petiolar bundles 
in Todea, as described by Faull (1901). In the very young 
bundle there are four to six proto-xylem groups. In the mature 
leaf the number is invariably four (Fig. 16) as in Osmundites 
skidegatensis. The bundle, however, is not concentric; as 1s 
the case in Osmundaceae (Faull); but the small celled phloem 
is situated at the dorsal side of the leaf, bending in at both 
ventral sides and terminating at the two inner proto xylems. 
The tracheae show numerous bordered pits. This bundle 
