TRYPANOSOMES IN TSETSE-FLIES AND OTHER DIPTERA. 171 
In one fly I noted oval, refringent, motionless bodies in the 
proctodeeum. In the sucking stomach I noted, in the wall, 
“Jone, greenish-yellow filaments, consisting each of a row of 
joints of varying lengths.””! 
Nomenclature of the structural parts of trypano- 
somes.—There is much diversity in the names applied by 
different authors to some parts of the trypanosome body, 
more especially with regard to the smaller chromatic nucleus, 
and also in the use of the terms anterior and posterior. ‘The 
smaller nucleus is termed usually in England the micro- 
nucleus; in France the centrosome; and in Germany the 
blepharoplast. Hach of these terms is open to objection, 
the term micronucleus, in my opinion, least of all, so long 
as it is used in a purely descriptive sense to mean simply a 
small nucleus; the danger is, however, that it leads to 
instituting a comparison with the micronucleus of Ciliata, 
which is a totally different structure, a reserve generative 
nucleus. Whatever the nature of the micronucleus of a try- 
panosome may be, there is no evidence that it is composed 
solely of generative chromatin. 
The terms blepharoplast and centrosome raise the whole 
question of the nature of the bodies so named—a question 
upon which it is possible to have much difference of opinion. 
It is enough for me to refer here to the recent memoir by 
Goldschmidt and Popoff (16), in which the subject is 
discussed at length, and to state my own views: I regard a 
centrosome as an achromatic body, in connection with a 
nucleus; and a blepharoplast as a body of the same nature 
as a centrosome, but in connection with a protoplasmic loco- 
motor apparatus, such as a flagellum or cilium. This is the 
sense in which I understand these terms; others may differ 
from me. 
With regard to trypanosomes, I may point out, in the first 
place, that the ‘ micronucleus” is certainly a chromatic body 
and cannot be classed with achromatic structures. Apart from 
its staining reactions, which are more intense than those of the 
1 Compare Stuhlmann (41, p. 47). 
