AZ E. A. MINCHIN. 
larger nucleus, I may cite the observations of Schaudinn (40), 
who describes it as arising in Trypanosoma noctne by an 
unequal division of the zygote nucleus; and I may further 
draw attention to the condition in Trypanoplasma, where 
it is as large as, or even larger than, the other nucleus, 
so that the term micronucleus for it becomes rather a 
misnomer. 
In the second place, I may point out, as others have done 
also, that the flagellum does not arise from the smaller 
nucleus, but quite independently of it, from a minute basal 
granule.! The behaviour of the flagellum and of the smaller 
nucleus in division also shows clearly their complete struc- 
tural independence, as I have pointed out below (p. 193). 
I consider the basal granule of the flagellum as a true 
blepharoplast in the sense in which I have defined the term 
above; and I regard the nuclear apparatus of trypanosomes 
as specialised into two distinct portions, one regulating the 
function of locomotion, the other that of nutrition. Hence I 
consider that the terms kinetonucleus and tropho- 
nucleus, suggested by Woodcock (42), express most 
correctly the true nature of these bodies, and I shall employ 
these terms in my descriptions, though it is often convenient 
to speak of the trophonucleus simply as the nucleus, and as a 
further abbreviation I shall sometimes refer to the two bodies 
as n and N simply. 
From the true blepharoplast arises the flagellum, which 
passes to the surface of the body and runs along the edge of 
the undulating membrane as the marginal flagellum, until it 
reaches the end of the body, where it becomes a free 
flagellum of greater or shorter length. According to 
Schaudinn’s observations on the formation of the locomotion 
apparatus in Trypanosoma noctue (40), there exists also 
a distal blepharoplast, as it may be termed, situated at the 
1 Dutton, Todd, and Hannington (15, p. 219) point out that “the thickened 
end of the undulating membrane ends not in it [#] but in a pinkish basal 
granule or ‘ diplosome’” (why “diplosome”? It is only double when about 
to divide). Compare Novy (33), pp. 5 and 6. 
