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duel ais fis 
SI) ju ZATION DES LÉ 
j up the Salles th divect combination of pallida violet and variegata — 
yellow in, any proportions, had no tendency to produce red, and my ‘ 
_ conclusion is that the mixture of these two pigments alone could i 
| never give a crimson. a 
As there are other yellow pigments, — soluble or sap yellow % 
pigments, — besides the more common plastid yellow, on the pos- 
sibility that the yellow of /lavescens was different from the yellow 
of variegata, crosses were made to test the effect of flayvescens yellow 7 
_in combination with other types. I had all the more expectation of — D 
finding it a reddening factor since from one of the earliest crosses 
made of Queen of May  Thorbeck a seedling appeared indistin- 
guishable from flavescens, which seemed evidence that flavescens > 
was in the parentage of Queen of May. The results have however been 
very conflicting. Some crosses with flavescens, when either a pli- 
cata or a squalens or an amena, or all three were also in the 
ancestry gave red or rose taned seedlings. Crosses with a variegata, 
flavescens X Maori-King gave some seedlings with a brighter 
colour in the falls than ordinary variegata varieties, viz. Glitter, but 
from the same seed pod came also blue-toned amæna, and lasty,from 
and cross flavescens X macrantha came one of the bluest-seedlings 
have raised. J Blue bird seemed, therefore, that the reddening effect 
was due to the presence of one of the other types in the parentage, = 
_and from other resulls, most probably plicata. But so far as my expe- 
riments have gone, it seems improbably that plicata in the ancestry Ne 
_ of Queen of May, for only one plicata has appeared out of 33 seedlings a 
_ from crosses of Queen of May by a plicata, and thatit ismost likely, aa 
an accidental self seedling. ie 
3° One of the earliest results observed from crosses of plicata pet. 
with almost all other types was the number of cases in which red a 
toned seedlings appeared. The explanation that first occurred to me, if 
_— that the plicata type might be due to two complementary fac- À 
_tors, one of which by itself — had a reddening effect, seems no 
longer tenable in view of later results, and I have no alternative 
theory to suggest except that possibly some other species such as Py 
| balkana in the ancestry of plicata. This is, however, a mere guess, “4 
| based chiefly on the distinction heard of plicata. This effect of pli- ae 
cata (in certain combinations, for there are notable exceptions) is, ‘ + 
however, | am sure, a fact, and a mere impression not as it is the be. 
| recorded result of very many diverse crosses. Among the old stan- ak. 
| dard varieties, also plicata, is in the parentage of dre — the red- 
dest red RTS and in that of Jacquesiana, the reddest of the a 
red squalens. And it can hardly be a coincidence that all my red- «#4 
dest toned seedlings have plicata in some degree in their ancestry | 
and all that have been tested have proved to be heterozygous for 
