84 NATIVE BRITISH ORCHIDACE^ 



Sub-tribe PHYSURINM 



Distinguished from the Spiranthinas by the pollinia built up of packets of pollen. 

 Mostly found in tropical Asia, a few in Africa and America. Only one European 

 species. 



Genus IV GOODYERA R. Br. 



Column short, horizontal. Stigma facing the lip, rostellum apical supported between 

 curved horns. Anther leaning forward over rostellum, to which the stalkless pollinia 

 become attached; pollinia built up of oval packets of pollen-tetrads. 



Small herbs with creeping rhizome, ovate stalked leaves and small flowers with 

 bag-like hypochile and recurved epichile, in a spike-like raceme. 



Goodyera resembles Epipactis in the shape of the lip, but its plan of construction 

 is essentially that of Spiranthes, which it closely resembles as far as the inflorescence 

 is concerned. It shows a remarkable evolutionary trend in the direction of the 

 Ophrydeje by the development of pollinia built up of packets of pollen tied together 

 by elastic threads, which coalesce at the apex of the polhnium into a flattened truncate 

 ribbon adhering to the back of the rostellum, but not forming a caudicle, except 

 in the foreign G. discolor, which has long apical caudicles.i The attachment of the 

 anther to a filament by its back resembles that of Cephalanthera. 



Goodyera R. Br., in Ait. Hort. Kew. ed. z, v, 197 (1813). Satyrium L. (1753). 

 Epipactis Boehm. (1760). 



I. Goodyera repens R. Br. 



PI. 13; PL H, fig. 2 (p. 219). Creeping Lady's Tresses 

 Creeping Goodyera 



Rhizome creeping, short-jointed, with white slender runners extending tlirough the 

 layer of pine-needles without entering the soil, often emitting other runners, and 

 ending in a rosette of leaves so that a plant may give rise to a number of others all 

 connected together. Roots few, short, thickly clothed with hairs, brown, descending. 

 Flowering stem 10-25 cm. tall, erect, stiff, cylindrical, ridged above (through decur- 

 rence of the nerves of the bracts), pale green, glandular-hairy, with a whitish veined 

 leafless sheath at the base, sometimes with a leaf-like tip. Basal leaves almost in a 

 rosette, ovate or ovate-lanceolate, obtuse or acute, firm, rigid, keeled, narrowing into 

 a winged stalk sheathing the stem, dark green often marbled with lighter green, 

 5 -nerved, conspicuously net-veined, persisting through the winter; upper leaves 



' Darwin, Feri. Orch. ed. 2, p. 105. 



