OPHKYDEjE 121 



divisions. The position of the flower with tlie lip uppermost finds its counterpart 

 in Liparis (Malaxideas) and Nigritella (Gymnadeni^). R. Brown placed it in the 

 Gastrodias, especially on account of the deciduous anther, but Rohrbach, who devoted 

 much study to the plant, maintains that the anther is persistent, in which Endlicher 

 concurs. It is often placed in the Neottieae, having an epichile and clinandrium as in 

 Epipactis, but not a cup-like hypochile. None of the Neottieas has inverted flowers 

 or a coralloid rhizome, however, nor is any genus constructed on the same plan 

 as Epipogon. The pollinia are built up of pollen-packets as in the Physurinas, but 

 there is no other resemblance to that sub-tribe. The coloured ridges of the Up are 

 suggestive of Cephalanthera, but nothing could be more different from the latter than 

 the pollinia, long caudicles, and large rostellum of Epipogon. 



Tribe IV OPHRYDEiE Lindl. 



Anther in one piece with column. Pollinia two, built up of packets of poUen-tetrads 

 attached by elastic threads, and affixed by caudicles in a few species to one viscidium, 

 but as a rule to two separate viscidia. 



Sub-tribe I GYMNADENliNiE Engler, i)///^^. (1892) 



Viscidia not enclosed in a pouch left behind when they are withdrawn. 

 Ebursiculat^ Rchb. £ 



Sub-tribe II SERAPiADiNm Engler 

 Viscidia enclosed in a pouch left behind when they are withdrawn. 



BuRSICULAT^ Rchb. f. 



This division is based on a single character, and is to a certain extent artificial, the 

 latter sub-tribe including the genera Anacamptis and Neotinea, which have the 5-lobed 

 column and separate lateral stigmas of the genus Gymnadenia. 



16 



