OPHRYDE— MGYMNADENIIN^— GYMN ADEN 1 A 



'39 



but the reproductive organs of P. hijolia. The hybrid is sotBetimes nearer one parent, 

 sometimes the other. 



It is recorded (/.i3. p. 95 (19 10)) that P. chlorantha occurred in Fhntshire with plenty 

 of P. bifolia, but that careful search revealed no intermediates. This was to be expected, 

 hybrids between the two being very rare. A specimen of P. chlorantha was exhibited 

 to the Linnean Society with all tliree petals spurred — a case of true peloria. A speci- 

 men figured in their Journal {Bot. xxxviii, t. i) had three sepals spurred — false peloria 

 — a really extraordinary accident. 



A single specimen of this hybrid found amongst multitudes of both parents at 

 Sligachan, Skye, differed from P. bifolia by its longer spur and slightly divergent 

 anther-cells, and from P. chlorantha by its colour, and the shape of the sepals and spur 

 {B.E.C. p. 508 (1910)). With it was found a peloriate specimen of P. chlorantha in 

 which the lip had reverted to a spurless petal. 



Sowerby (£.B.) states that he found specimens on Reigate hills difficult to assign to 

 P. bijolia or chlorantha, but that he did not then know of the differences in the pollinia, 

 caudicles, viscid discs, and stigmas. 



Genus XV GYMNADENIA R. Br. 



Column short with an oval lobe on each side, on the front of which the stigmas are 

 situated. Pollinia erect on forward ends of the two parallel linear viscid glands. 



Herbs with palmate or fascicled tubers, long narrow or ovate-oblong leaves and 

 cylindrical spikes of small flowers with spreading side-sepals, petals connivent with 

 upper sepal, short 3-lobed lip, and spur with free honey. 



About 12 known species, of which four inhabit Europe, and the rest Central and 

 Eastern Asia to Japan. Gymnadenia R. Br. in Ait. Hort. Kew. ed. 2, v, 191 (18 13) was 

 included in Or^/;/.f by Linnseus {Sp. pi. ed. i, p. 942 (175 5)) and in Habenaria by Bentham 

 and Hooker. The latter genus is now restricted to exotic orchids with projecting 

 stigmatic processes, and has no European representative. A few authors still cling to 

 the genus Habenaria^z heterogeneous collection of widely diifering genera, including 

 Gymnadenia, Platanthera, Caloglossum and Neotinea. 



Meyeri created a new genus, heucorchis, for G. albida, the following being a trans- 

 lation of his diagnosis: "Lip through connivence with the other perianth-segments 

 almost campanulate. Each pollen-mass attached erect to a separate, naked gland. 

 Everything else as in Gjmnadenia" . As each pollinium is similarly attached to a naked 

 gland in Gjmnadenia, Meyer's new genus is only based on the sepals, petals and lip 



' Preuss. Pflam^engatt. (1839). 



18-2 



