OPHKYDEMSEKAPIADINJE 151 



There are two anomalous genera placed under this sub-tribe because the pollinia 

 are enclosed in a common pouch — Anacamptis and Neotinea. They both have a 

 3-lobed column with two separate stigmas situated on the side-lobes, as in Gymna- 

 denia, instead of the central compound stigma on the roof of the spur-entrance as 

 m Orchis. Anacamptis is really a Gymnadenia as far as the sepals, petals, lip, spur, 

 and 3-lobed column with lateral stigmas are concerned. It has, however, taken three 

 steps forward in evolution in the direction of Orchis, (i) The linear horizontal back- 

 ward pointing viscid glands of Gymnadenia have taken a half-turn outwards, and the 

 broader ends have united, forming a transverse strap-shaped viscidium with the erect 

 pollinia rather close together in the middle. (2) It has evolved a pouch like that 

 oi Orchis, to keep the viscid gland moist and sticky. (3) It has ceased to provide free 

 nectar in the spur, and secretes liquid instead between the spur-waUs as in Orchis. 



Neotinea resembles G. albida in the one-sided spike of small almost tubvdar (usually) 

 white flowers with connivent sepals and petals, 3-lobed lip, short honey-filled spur, 

 lateral divergent stigmas on side-lobes of column, and very smaU pollinia with rela- 

 tively large rounded pollen-packets and extremely short caudicles. It has, however, 

 evolved the rounded entire tubers, and the orbicular viscidia enclosed in a common 

 pouch, of Orchis. 



KEY TO THE GENERA 



A. Both pollinia attached to a single viscidium enclosed in a pouch {bursiculd). 



1. Flower very like Gymnadenia conopsea, long-spurred, but with two erect guiding 



plates on base of lip. Viscidium strap-shaped, the ends curling round and 

 clasping the proboscis of Lepidoptera. Anacamptis 



2. Flowers green with long narrow ribbon-like lip and short spur. Viscidium large, 



flat. Himantoglossum 



B. Each pollinium attached to a separate viscidium. 



3. Flower without spur. Viscidia sometimes accidentally coherent (but often removed 



singly by insects). Aceras 



4. Flower spurred, without free nectar. Orchis 



5. Side-sepals saccate at base. Spur very short, nectariferous. Stigmas two, on 



side-lobes of column. Neotinea 



6. No spur. Each viscidium in its own separate pouch. Stigmas confluent on inner 



surface of chamber forming foot of column. Ophrys 



