ON THE STRUCTURE OF JINIGMA iENIGMATICA. 289 



prises, et sur d'excellentes preparations au carmin alune, j'ai 

 pu la suivre aussi bien aa niveau de I'epithelium modifie que 

 sur la reste de la section." I think Barrois must have been 

 mistaken on this point, but however that may be, the modified 

 cilia are very conspicuous in ^Enigma. Their arrangement 

 leaves no doubt that the groove has a function analogous 

 to that of the endostyle of an Ascidian, and that its cilia 

 sweep a current of liquid or viscous matter (I could find no 

 solid particles in it) along the length of the ceecum. 



From a consideration of these histological details, I suggest 

 that the material of the crystalline style is secreted by the 

 yellow glandular cells of the epithelium of the stomach, and 

 that the so-called cuticular lining of the stomach, or ^'fleche 

 tricuspide " of Poll, is nothing more than the coagulated 

 viscous secretion of these glands. This viscous secretion, 

 I suggest, is swept into the caecum by the action of the cilia 

 of the ciliated groove, and is there moulded and solidified into 

 the substance of the crystalline style which, as Mitra has 

 proved, is a globulin containing an amylolytic ferment. During 

 the process of digestion, the style as a whole is moved forward 

 into the cavity of the stomach by the action of the stiff brush- 

 like cilia of the normal caecal epithelium, and is gradually dis- 

 solved, liberating the amylolytic ferment. It is true that 

 several authors, including Barrois, have asserted that the 

 anterior end of the crystalline style is continuous with the 

 so-called '^fleche tricuspide," but this confirms rather than 

 contradicts my suggestion. These same authors assert, and 

 I agree with them, that the substance forming this cuticular 

 coat or "fleche tricuspide" is identical with the crystalline 

 style. When one examines this substance carefully, one 

 finds that it forms a lining to the wall of the stomach, thin in 

 some places, thicker in others, and where it is thicker its 

 inner surface (that is the surface farthest from the epithelium) 

 passes insensibly into a mass of granular coagulum, in which 

 the anterior end of the style appears in some cases to be 

 imbedded. But there is no real continuity between the two. 

 The anterior end of the style is in all cases much reduced in 



