152 



PROCEEDINGS OF THE NATIONAL MUSEUM 



Bradycinetulus Cockerell 



B. fossatus (Haldeman) 



B. ferrugineus (Palisot de Beauvois) 



B. rex Cartwright 

 Bolborhombus Cartwright 



B. carinatus (Schaeffer) 



B. angulus (Robinson) 



B. parvulus Cartwright 

 Bolbocerastes Cartwright 



B. regalis Cartwright 



B. imperialis Cartwright 



B. imperialis kansanus Cartwright 



B. serratus (LeConte) 



B. peninsularis (Schaeflfer) 

 Bolbelasmus Boucomont 



B. arcuatus (Bates) 



B. minor (Linell) 



B. hornii (Rivers) 

 Eucanthxis Westwood 



E. lazarus (Fabricius) 



E. lazarus subtropicus, new subspecies 



E. alutaceus Cartwright 



E. greeni Robinson 

 Bolboceras Kirby 



B. obesus (LeConte) 



B. falli (Wallis) 



B. thoracicornis (Wallis) 



B. cornigerus Melsheimer 



B. Uebecki (Wallis) 



B. filicornis (Say) 



B. floridensis (Wallis) 



B. simi (Wallis) 



B. darlingloni (Wallis) 



B. alabamensis (Wallis) 

 Geotrupes Latreille 



G. opacus Haldeman 



G. ulkei Blanchard 



G. blackburnii (Fabricius) 



G. blackburnii excrementi Say 



G. egeriei Germar 



G. hornii Blanchard 



G. balyi Jekel 



G. stercorarius (Linnaeus) 



G. splendidus (Fabricius) 



G. splendidus miarophagus Say 



G. semiopacus Jekel 

 Peltotrupes Blanchard 



P. profundus Howden 



P. profundus dubius, new subspecies 



P. youngi, new species 

 Mycotrupes LeConte 



M. retusus LeConte 



M. lethroides Westwood 



M. gaigei Olson and Hubbell 



M. cartwrighti Olson and Hubbell 



M. pedester Howden 



In the discussion, an attempt is made to compare the habits of North 

 American Geotrupinae with the habits of Geotrupinae from other 

 continents, particularly Europe. The feasibility of using food and 

 burrowing habits as aids in taxonomic and phylogenetic investigations 

 are discussed, and one case is pointed out where they could be applied. 



ACKNOWLEDGMENTS 



Much of the work has been carried out at North Carolina State 

 College under the able direction of Dr. P. O. Ritcher, Oregon State 

 College, who aided me greatly in the field, taking many of the photo- 

 graphs included here and making many helpful suggestions during 

 the preparation of the manuscript. I am also extremely grateful to 

 O. L. Cartwright of the U. S. National Museum for aid in checking 

 identifications and for many helpful suggestions on the taxonomic 

 aspects of the work. 



Special thanks are also due to Dr. Henry K. Townes, North Caro- 

 lina State College, for his help both in the field and on the manuscript, 

 and to Dr. T. H. Hubbell, University of Michigan, whose comments 

 and manuscript enabled me to write the summary on Mycotrupes. 



