21 
Liophis oligolepis Boulenger 1905:455. Type locality: Igapé-Assu, Para, 
Brasil. 
Leimadophis reginae macrosoma Amaral 1935:238. Type locality: Canna Brava, 
Leimadophis reginae maculicauda Hoge 1954:241. 
Goias, Brasil. 
Leimadophis zweifeli Roze 1959. Type locality: Rancho Grande, Aragua, 
Venezuela. 
Liophis reginae, Dixon 1980:24. 
DISTRIBUTION: Cis-Andean South America, from Colombia to northern Argentina; 
also Trinidad and Tobago (see figure 2, p. 118, Dixon 1983). 
SUBSPECIES: Four. 
Ihe 
a 
b 
Dorsum with small black and yellow spots; black lateral 
Calidal Stripe faint Of abSeENt 22. cee emcee] ce =< le gees 
Dorsum greenish, olive, or grayish, never with small 
yellow and black spots; black lateral caudal stripe 
always wresent tand Wistinctva2k. 2ytetejonloi: Sines Joga. 233 
Subcaudals average 80 (69-88) .................. reginae zweifeli 
Subcaudals average 65 (55-78) .............. reginae semilineatus 
Dorsal scale rows one and two pale colored,in contrast 
to.dorsall coloration, [6:340!. tack 20. eres Bayo. fans. et: 
Dorsal scale rows one and two similar in color to rest 
Of bodyveehoretahens bre oeeee, «2st Leelee rp yao. 100. at 
Dorsum with dense pale and dark paravertebral flecking; 
subcaudals average 74 (63-80) .......-..5...-. reginae reginae 
Dorsum without pale and dark paravertebral flecking; 
subcaudals average 67 (63-80) ........... reginae semilineatus 
Subcaudals with ventrolateral black spots, flecks, or 
smudges; subcaudals average 81 (75-91) . reginae macrosomus 
Subcaudals imseniliates subcaudals average 70 (55- 81) 
sis ett + + sSBeUEG Le eekh -PEBReGEeGhEeu menor reginae semilineatus 
Liophis reginae reginae (Linnaeus) 1758. 
SYNONYMS: violaceus Lacépéde; graphicus Shaw. 
DISTRIBUTION: Guyana, Suriname, and French Guiana (see figure 2, p. 
Dixon 1983). 
Liophis reginae macrosomus (Amaral) 1935. 
SYNONYM: maculicaudus Hoge. 
DISTRIBUTION: Chaco and Cerrado of Argentina, Bolivia, Brasil, and 
Paraguay (see figure 2, p. 118, Dixon 1983). 
Type locality: none given. 
118, 
