Phylogeny of the Pierines. 293 
to are (1) the streaks and touches of yellow that begin 
to occupy much of the area of the original pale patches, 
including, as has been seen, the spots of I on the hind- 
wing; and (2) the bright red patches visible in the 
basal region of the same wing close to the body. These 
have been already discussed at some length (see 
pp. 281, 285), and their importance has been shown in 
reference to the markings of more recent groups. In 
this place it is only necessary to note once more that 
their ancestral character seems to be proved not merely 
by their existence in a fully developed condition in 
genera so widely separated geographically as _ the 
Western Catasticta and Leodonta and the Hastern 
Prioveris and Delias, but also by the fact that there is 
scarcely a genus throughout the whole sub-family, 
whether in the Old or New World, that does not show 
some relic of their former presence. With regard to 
their origin, no clue appears now to exist. It seems 
impossible to trace them further back than to the 
EKastern and Western genera named; and the probably 
still more primitive form, Huchetra socialis, to which we 
should naturally turn for an indication of their develop- 
ment, affords us in this particular no information 
whatever. — 
In the genus Leodonta we find some modification in 
the shape of the wings, and only three instead of four 
branches to the subcostal nervure of the primaries. But 
the wing pattern remains much the same as in those 
species of Catasticta where the white central band has 
grown into a well-defined area and the two series 8 and 
M have not yet become clearly distinguishable. The 
yellow streaks and red basal marks are still prominent 
on the hindwing beneath, and the separation between 
S and M is, in this genus also, more evident on the 
underside. 
In Pereute and Huterpe, which are undoubtedly very 
close allies of the genera just referred to, the development 
of the pattern has taken a somewhat different direction. 
The primitive dark ground colour is in the males of 
several species of Pereute, as P. charops, P. autodyca, and 
the Hewitson collection labelled as C. notha. This is undoubtedly 
erroneous, as the underside of the true C. notha is almost exactly 
like that of C. corcyra. 
