310 Professor KE. B. Poulton on Mimetic Forms of 
the National Collection. In view of these intermediate 
specimens, and the variation in all the distinguishing char- 
acters observed when a sufficiently long series of johnstont 
are examined I do not doubt that Aurivillius is correct 
in suggesting that fallax is conspecific with johnstoni. 
Strong support is also afforded to Aurivillius’ suggestion 
by the observations of Rev. K. St. Aubyn Rogers, who 
knows both johnstont and fallax in life in their natural 
habitat and looks upon them as a single species. It has 
been shown here that fallaa is undoubtedly the eastern 
form of lycoa. It therefore becomes extremely probable 
that the whole wonderful series of forms—many of them 
totally unlike—associated under the name johnstoni, or 
as it was still more appropriately named by Oberthiir, 
proteina, are all of them specifically identical with Godart’s 
species lycoa. Furthermore, this remarkable series must 
be still further extended to include the torwna of Grose- 
Smith. 
In conclusion, it is possible to attempt to reconstruct 
the history of the changes through which /ycoa and its 
descendants have passed. It is probable that the male 
of the western lycoa represents the ancestral form of the 
whole group,—a semi-transparent fuscous and brownish 
Acrew with ill-defined markings. As regards the semi- 
transparency it is noteworthy that the character tends to 
crop up not uncommonly in the most modified form 
Johnstont, where it is often seen in the discal patch of 
the hind-wing. The female of the western lycoa became 
modified by synaposematic approach to the black and 
white species of the Danaine genus Amawris on the 
west coast. ‘The same is substantially true of the species 
in Western Uganda where the black and white Amauwris 
are still predominant and have even drawn the echeria and 
allimaculata types of their own genus after them. (See 
S. A. Neave in Trans. Ent. Soc. 1906, pp. 208-210.) As we 
go further east however these latter types become them- 
selves predominant, and the fallax forms of lycoa follow 
them, the males becoming strongly mimetic and approach- 
ing the buff: spotted Danaine models, while the females 
still retain the ancestral colour and resemble those that 
are white-spotted. As regards the hind-wing both sexes 
gain a buff discal patch similar in colour but not in shape 
to the models. Finally, from the most strongly-marked 
of these eastern forms with the deepest shade of ground 
