3!58 j\Ii'. R. I. Pocock on some of ihe External 



its way to the exterior along a narrow passage — tlie tliree 

 jjassages converging and fusing to forni a common duct 

 opeuiug to the exterior witliin the anal pouch. In the 

 exauiple of Hycena hycena, on tiie contrary, the normal an.d 

 gland is piriform and saccular and nndivided, and opens at 

 its narrow end by a small duct into the anal pouch. Close 

 to its narrow end, externally and in front, there is a large, 

 double, supplementary, glandular mass, which differs from 

 the fiask-shaped or piriform gland in containing no cavity 

 and no definite duct. The secretion from this gland makes 

 its way into the anal ponch by means of a number of minute 

 orifices scattered over the wall of the pouch adjacent to the 

 orifice of the duct of the piriform gland, which unmistakably 

 corres[)onds to the normal anal gland of other Carnivora. 



According to Murie, therefore, H. hrunnea has a single 

 pair of anal glands, each subdivided into three compart- 

 ments, the passages from which join to form a common duct, 

 and there are no accessory glands ; whereas in H. Iiyeena 

 there is a single pair of simple undivided anal glands, each 

 being accompanied by an accessory mass of enlarged cuta- 

 neous glands opening into the anal pouch by numerous 

 small apertures. Consideiing the tolerably close resem- 

 blance in other respects between the two species, this 

 difference is full of interest. 



Judging from Watson's account of these glands in 

 Cvocuta crocuta (P. Z. S. 1877, p. 369, pi. xli. and 1878, 

 )). 410, pi. XXV.), there is also a single pair of piriform anal 

 glands in the Spotted Hyjiena, and these are connected along 

 the lower portion of the pouch by a band of accessory glands 

 opening into the pouch by a " line of perforations." 



The very exact and detailed account given by Daubenton 

 (Buffon's Hist. Nat. ix. pp. 287-288, pis. xxvii. & xxviii. 

 1761) of the glands in the Striped Hyaena agrees closely 

 with my observations upon that species, exce})t that in 

 Daubenton's example the walls of the sack above the rectum 

 Avere more highly glandular than in my example, and the 

 lateral glandular mass does not appear to have been in any 

 way subdivided. 



Mivart (P.Z. S. 1882, pp. 198-199 and 201) summarised 

 the facts recorded by Daubenton, Murie, and Watson by 

 saying " There is an anal pouch with two {H. str'iata^= hyaena) 

 or tliree (H. brunnea) pairs of anal glands on each side of the 

 rectum; and in one [i7. hy(sna~\,'\i not in both, species 

 there is a transverse band of isolated [glandular] follicles 

 at the bottom of the anal pouch " (pp. 198-199) ; and in 

 Crocuta '•' there is but a single pair of anal glands, one on 



