430 On the Anatomy of Paludestrina jenkinsi. 
in the same tubules as ripening odcytes and also in the lining 
epithelia of such tubules (Pl. XV. figs. 7A & B). 
The oviduct, which passes down the columella area in 
company with the genital artery, is exceedingly slender and 
usually thin-walled. At its distal end it becomes rather con- 
voluted and thicker. It opens into the vagina very close to 
and probably in association with the spermatheca. ‘The latter 
is an organ corresponding in shape and position with the 
spermatheca of normal female gastropods, and there can be 
little doubt that it represents that organ. In the present 
instance, of course, it is functionless. The persistence of 
such an organ ina parthenogenetic female is very interesting, 
and not without importance in relation to the general question 
of pepe lag A spermatheca is found in a few other 
Teenioglossa, e. g., Cyclostoma (4) and Paludina (3). In other 
genera riatalipuowidened close to Paludestrina there is noacces- 
sory organ in the female genital complex ; but there appears to 
be some evidence for its presence in P. ulve (see below). 
The albumen-gland is a large organ readily observable in 
the living animal lying at about the junction between the 
fifth and sixth whorls, It opens into the brood-pouch by a 
vertical slit-like aperture adjacent to the commencement of 
the vagina. 
The vagina is very difficult to locate exactly, owing to 
contraction ; it appears to be a tube passing forwards beneath 
the brood-pouch on the right-hand side, and opening into the 
mantle-cavity near the anus. The communication between 
the vagina and brood-pouch would appear to be a fairly wide 
aperture rather anterior to the openings of the albumen- 
gland and oviduct (Pl. XV. fig. 8). 
The brood-pouch is a very capacious cavity when fully 
extended, It is excavated in the pallial integument of the 
right-hand side. On the left it is bounded by the rectum. 
When functional it occupies the whole right-hand side of the 
last whorl and almost reaches the penultimate whorl. In 
this condition it is capable of holding well over forty young. 
Lehmann (6) has described the male and female genitalia of 
- H. balthica (=P. ulve), but neither his description nor his 
illustrations are very satisfactory. If his account is to be 
followed, the brood-pouch leads on into the vagina, which is 
not the condition seen in jenkinsé. On the other hand, he 
says: “ (die Scheide)... von welcher ein kurzer Blasenstiel 
nuit rundlicher Blase ausgeht,” which seems to agree with the 
spermatleca described above for yenkinst. He also figures 
and describes an apparently distinct and well-marked 
albumen-gland. 
It should be pointed out that among other Tenioglossa 
