38 Prof. Link on the Structure of the Orchidacee. 
among which one on each side might readily be supposed to 
indicate a stamen on each side. The other canal, which is there 
represented, originates from the confluence of the labellum with 
the column, and whenever fertilization was artificially effected, I 
found poilen-tubes in this canal also. Similar transverse sections 
of the column of other Vandee always exhibit a quantity of vas- 
cular bundles surrounding the stigmatic canal. It is clear there- 
fore that there is still another part which surrounds the column, 
and with the labellum represents a special organ, which must be 
referred to the Linnean nectary or to an accessory corolla (para- 
corolla). This accessory corolla has two lips; one, the upper lip, 
blended with the column, the lower lip being the labellum. 
The comparison of the Orchidacee with the Alpiniacez lies 
near, and indeed has been occasionally made, although mostly 
superficially. The calyx of the Alpiniacee is spathaceously tri- 
foliate, and, according to Lindley, corresponds to the calyx of 
the Orchidacez : the corolla of the Alpiniacez always has two 
divisions ; the outer tripartite envelope can only be compared, ac- 
cording to Lindley, with the corolla of the Orchidacee, where 
however that third leaflet is wanting, being absorbed, as it were, 
into the labellum situated above it. There is nothing in the 
flower of the Orchidacez corresponding to the inner division of 
the flower of the Alpiniacez, unless the envelope of the column, 
above spoken of, is taken into consideration. This, together with 
the labellum, corresponds to the imner portion of the flower of 
the Alpiniaceze, in which there is always a well-marked labellum, 
and very. often, for instance in Hedychium and Globba, an upper 
lip, which is merely not blended with the stamens and style as 
in the Vandee. This upper lip is often wanting and the label- 
tum exists alone, as in Alpinia, Zingiber and Kempferia, just as 
in our indigenous Ophrydee. The connecticulum is very much 
expanded in the Alpiniacez, and so it is in the common anthers 
of the Ophrydee, as well as in the calyptrate anthers of the 
Vandee, and indeed in all Orchidaceze the two anther-cells are 
connected above by a membranous or fleshy portion, which may 
be aptly named a connecticulum. 
As to the anthers, I will merely observe that the pollen of the 
Vandee does not always lie naked upon the cellular body which 
serves as its basis, but is inclosed in a delicate membrane of an- 
gular parenchymatous cells, as I have distinctly seen in many, 
particularly in Huntleya violacea. 
Cypripedium is not diandrous. The column divides into two 
branches, each of which bears an anther-cell with two pollen- 
masses. This division of the column alone distinguishes this 
genus from the rest. Only one anther exists, but its chambers 
are very much separated, as is usual in the Alpimacee. My re- 
