External Characters of Ruminant Artiodactyla. 441 



well developed as in the male. Their secretion had a 

 decidedly pungent and unpleasantly musteline odour. 



The rhinarium is well developed and " bovine." From 

 the anterior aspect the upper margin is strongly convex and 

 the area beneath the nostrils is mesially grooved and very 

 wide — wider, in fact, than the area above those orifices — and 

 visible to a considerable extent in profile view. From the 

 dorsal sidte the anterior margin is convexly truncated, and 

 the posterior margin is straight between the posterior angles 

 of the nostrils, the hair of the nose not extending forwards 

 beyond that line. 



Genus Boskt.aphus, Blainv. 

 Boselaplius tragocamelus, Pall. (p. 9?(3). 



In a male example the preorbital gland had a much 

 shallower pit than in the female described in 1910, and was 

 without definite lids. The gland itself, moreover, was not 

 regularly heart-shaped, but was longer than thick and of 

 irregular form. 



The rhinarium (fig. 1, A, B, C) is large and "bovine," 

 closely resembling that of Tetraceros, but more protuberant 

 in front, and, beneath the nostrils, laterally and with a 

 wider internarial septum. On its dorsal side the hair 

 advances a little way between the nostrils, so that the poste- 

 rior border of the rhinarium is concave. 



In 1910 I briefly described the glandular nature of the 

 skin between the ialse hoofs of the hind feet in the female. 

 The same feature is present in the male where the skin 

 between the widely separated false hools is clothed with 

 longish hair, is very thick and glandular, and mesially 

 lolded. In the fore foot there is no trace of the gland, the 

 false hoofs being larger and the hair restricted to the narrow 

 area between them. This gland (fig. 3, B) on the hind foot 

 of Boselaplius clearly represents an earlier stage of the 

 evolution of the pair of pouch-like glands present in Tetra- 

 ceros. The presence of similar glands in Taurotragus and 

 Strepsiceros (cf. infra) serves to link Boselaplms with the 

 African Tragelaphines, and refutes, if refutation be needed, 

 Kutimeyer's claim that Boselaplius belongs to a different 

 group. 



Inguinal glands are absent and there are two pairs of 

 mamma. 



The penis (fig. 1, I), E) agrees, generally speaking, with 

 the sketch and description published by Gerliardt (pp. cit. 



Ann. & Mag. N. Bist. Ser. i). Vol. ii. 32 



