450 Mr. R. I. Pocock on some 



alleged genera and subgenera. Since that year, however, 

 study of certain other external features — notably the rhina- 

 rium and penis — have supplied additional characters to those 

 derived from the skull, horns, tail, distribution of hair, and 

 outward form, which, I think, justify Gray's claim that the 

 groups are worthy of generic recognition. Probably other 

 characters bearing out this view will come to light with the 

 examination of further material. 



So far as the cutaneous glands are concerned, the genera 

 have the following mainly negative features in common : — 



Preorbital glands, as in all African Tragelaphines, are 

 absent. 



Inguinal glands are invariably absent, as in the Trage- 

 laphiue genera Taurotragus, Boselaphus, and Tetraceros. 



Pedal glands of the interdigital lype are also invariably 

 absent, as in all Tragelaphines. 



Glands on the false hoojs are absent, as in Tragelaphus. 



Two pairs of mammae, are present, as in all Tragelaphines. 



Genus Bos, Linn. 



Bos, Linn. Syst. Nat. ed. 10, p. 1758 : typo, taurus. 



Rhinarium (figs. 1, A, B ; 3, C) large ; viewed from the 

 front its upper margin is evenly convex from side to side 

 and the median area below the line of the widely separated 

 expanded nostrils is wider than the internarial septum 

 throughout its extent, the hairs of the upper lip extending 

 inwards neither beneath the nostrils above nor along the 

 edge of the upper lip below ; above the edge of the lip there 

 runs upwards a short shallow median groove, which is 

 present in all genera, and thus disproves Lydekker's state- 

 ment (Cat. Ung. in Brit. Mus. i. p. 11, 1912) that the 

 rhinarium in the Bovinse is undivided. A few scattered 

 hairs arise from the rhinarium interiorly, and its surface is 

 sculptured and reticulated. The anterior portion of its 

 dorsal surface is exposed to a varying degree in accordance 

 with the extent to which the hair of the upper side of the 

 muzzle spreads forwards between the nostrils; but the naked 

 upper edge of the nostrils is always of considerable width 

 and depth, and not narrowed as in Bison and Poephagus. 

 The extension of the hair between the nostrils above varies 

 according to the breed, being greater, for instance, in British 

 park cattle (B. taurus) than in Indian humped cattle (B. in- 

 dicus) ; but intergradation between these two forms seems 

 to be supplied by other breeds of B. taurus. 



