REVISION OF THE NATURAL ORDER HEDERACEJ. 231 
has been worked up, I will give a proper systematic arrangement of 
them. At present there is not even a complete list of the genera. 
I. ON THE POLYANDROUS GENERA. 
In the * Botanical Magazine’ for April, 1856, there is published, on 
plate 4908, a new genus of Hederacee, Tupidanthus calyptratus, Hook. 
fil. et Thoms., of which it is remarked, that **the coalescence of the 
calyx lobes and corolla into an arched coriaceous calyptra, together 
with the numerous stamens, the total absence of styles, and very 
numerous cells of the ovary, are perhaps unique in the Order” to 
which it belongs. The characters are certainly very singular, but it had 
evidently been overlooked that two years previously (in 1854) Asa Gray 
described two allied genera from the South Sea Islands (Plerandra and 
Tetraplasandra), both of which share with Tupidanthus a calyptrate co- 
rolla, polyandrous stamens, and a many-celled ovary. The calyptrate 
corolla had previously been noticed in some of the older genera of the 
Order, but the polyand t we tainly quite a new feature,— 
no more 13 than had been known to exist amongst this group of plants. 
During my exploration of the Viti Islands, I was fortunate enough to dis- 
cover several additions to the polyandrous Hederacee, among them two 
entirely new genera (Nesopanar and Bakeria), and in the following 
ages I propose to give a description of them, together with an enu- 
meration of all the polyandrous Hederacee known to me. In this list 
a polyandrous genus will be missed, which being referred by Bentham 
and Hooker fil. (Gen. Plant. p. 17) to Hederaceea,—* est Araliacea 
anomala ovario subsupero ” are their exact words,—would naturally be 
looked for in this place. Imean Zrochodendron, a Japan genus founded 
by Siebold and-Zuccarini, and placed by them amongst Winterea, by 
Endlicher as an anomalous genus at the end of Magnoliacee, and by 
Miers in the neighbourhood of Ternstreemiacem. However it cannot 
be admitted amongst Zederacez, differing as it does from all the 
known members of the Order by its 4-celled anthers and many-ovuled 
ovaries, to say nothing of its entire want of calyx and corolla. Its affini- 
ties are, in my opinion, much more with JFinterez, the very group in 
which Siebold and. Zuccarini placed it; and its nearest ally I hold to 
be Euptelia, Sieb. and Zucc., the nature of which has become better 
understood by the recent publication of Drs. Hooker and Thomson’s 
new Indian species (‘Linnean Journal,’ sect. Botany, vii. 241. t. 2), 
