94 ON THE SEXUAL ORGANS OF THE CYCADACEJE. 
tissue is slightly different, but there is no definite membrane admitting 
of separation. I cannotel early comprehend what Gottsche means by the 
yellow membrane which, according to him, covers the endosperm (Bot. 
Zeit. l. c. p. 398), unless he has in view the remains of the nucleus. 
It is not unusual to see germination commence in the undetached 
seed itself ; the radicle pierces the remains of the summit of the nucleus 
and appears externally (Plate XCII. fig. 11). We must interpret in 
this way the figure of the embryo of Maerozomia which I published in 
1845 (Ann. des Sc. Nat. l. c.). In this instance the embryonic con- 
dition had been passed and the first leaves developed. 
It must be remarked that in all Cycads the plumule is only com- 
osed of a few scales. On this and other points I may refer to my 
communications in the Ann. des Sc. Nat. |. c., and in * Linnea’ xix. 
p. 5. The parts which produce the pollen,* and which compose 
the male cone, are in every point the morphological equivalents of the 
carpophylls. It would be perfectly correct to call them anthers, but 
from their analogy to the female organs, and to avoid the risk of con- 
founding them with their loculi (which many authors continue very 
improperly to call anthers), it will be perhaps preferable to adopt the 
term androphylls. They are always smaller and simpler in shape than 
the earpophylls, but their structure presents no essential difference. 
The polygonal cells of the epidermis are much thickened, and the sto- 
mata are deeply situated among them. 
The principal point which arises here relates to the development of 
the numerous polliniferous loculi (‘ Zogettes’) which are situated on 
the under surface of the androphylls, and which were formerly looked 
upon as distinct anthers.t Mohl has observed with reason that the 
way in which I had explained in my monograph the evolution and 
morphological meaning of these loculi was not consistent (* Vermisehte 
Schriften,’ p. 57). 
They are placed on the lower face of the androphyll (which is also 
their situation in the peltate forms) and on either side of the median 
liue, and they spring in groups of two to four, or rarely of five from a 
* The oe figures which exist of these organs may be con nsulted. 
T This was the opinion of Richard (Dict. Class. d'Hist. Nat. tome v. p- 
216). Other Hii s as to the d ps _ oo R. Brown in the Ap- 
pasi to Captain King’s as Bot. Wo bin vol. i. p- 
459, et seq.). Lindley eco, Wired. p.293), adoptin beti s view, terms 
the androphylls * antheriferous cone scales" —W 
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