3 
4 $ 
Be 3 © 
a * & 
o aD i] le) 
°, (a Pw) S 
B = a o S 
ie) oO fo) wn » 
% c 2 g e 
2 se) ag ° 
Oe 
y 
> 
~ 
se So 
% 2 : 
Ds ) 
7 4 
Sos Sx 
2 
&p 
Yo, 
amoenus 
group 
fulyus group 
lineolatus group 
D4 constrictus group 
scapalis group 
discoideus group 
profusus group 
algae ees 
quadrifer group get Sellen ea ay 
. instabilis group 
cinereus group 
qa aeniceeaey sculpticollis group 
~ Hedychrous ? 
oe (India) 
Afrosmicronyx ? =. io - 
(West Africa) = SS 
_-= lopelatus ? 
(Madagascar) 
Hypothetical 
Smicronychine Ancestor. 
Diacram 1.—The apparent phylogenetic relationships among the groups of species of 
Smicronyx of America north of Mexico. 
The three important general changes or “‘steps” 
referred to in the text are indicated by Roman numerals 1, 1, and 11. 
Capital letters 
indicate subgenera as follows: A, Smicronyx; B, Pachyphanes; C, Pseudosmicronyx; D, 
Desmoris. 
