450 PROCEEDINGS OF THE NATIONAL MUSEUM VOL. 118 
The subspecies is limited to the Pilcomayo River area, on the 
Argentina-Bolivia border; all specimens of torallyx from north of 
that area belong to different subspecies. 
Peronaeus (Lissoacme) longinquus (Morelet) 
Bulimus longinquus Morelet, Series conchyliologiques, vol. 3, p. 195, 1863. 
Drymaeus longinquus, Pilsbry, Manual of conchology, ser. 2, vol. 11, p. 293, 1898. 
Type locality: Western slope of Vilcanota range, southeastern Peru. 
One specimen (USNM 250244) from southern Peru, rare. 
Genus Paeniscutalus Wurtz, 1947 
Paeniscutalus crenellus (Phillippi) 
Bulimus crenellus Philippi, Malakozool. Blatt., vol. 14, p. 67, 1867. 
Megalobulimus (Microborus) incarum Pilsbry, Nautilus, vol. 58, p. 29, 1944. 
Bulimulus (Paeniscutalus) incarum Wurtz, Nautilus, vol. 61, p. 12, 1947. 
Type locality: ““Hacienda de Unigambal”’; Huaraz, Peru, for 
incarum. 
According to the lots studied in the U.S. National Museum and 
the Museum of Comparative Zoology and according to Bequaert’s 
opinion (Bull. Mus. Comp. Zool., vol. 100, p. 169, 1948), ancarum 
Pilsbry is a synonym of crenellus Philippi. The principal differences 
of Paeniscutalus are anatomical; the protoconch of the shell shows 
axial ribs broken or segemented into elongated granules. Wurtz 
stated that the species, by its conchological characters, is placed “‘in 
the second division of Bulimulus (sensu lato)”; inasmuch as these 
Bulimuli are now separated into several genera, very different from 
Bulimulus, the position of Paeniscutalus should be that of a genus 
rather than a subgenus. 
Genus Scutalus Albers, 1850 
Scutalus tupacii (d’Orbigny) 
Helix tupacii d’Orbigny, Mag. Zool., vol. 5, p. 16, 1835. 
Bulimulus (Scutalus) tupacit, Pilsbry, Manual of conchology, vol. 11, p. 19, 1897. 
Scutalus tupacii, Parodiz, Nautilus, vol. 70, p. 134, 1957. 
Type locality: La Paz, Bolivia. 
This is the only species of the genus found as far south as Argentina, 
and it is one of the most characteristic and abundant forms in the 
northwestern part of this country, in the provinces of Salta and 
Jujuy, and especially in Tucum4n; it also extends a little to the east, 
in Santiago del Estero. Besides many lots in the U.S. National 
Museum, the Museum of Comparative Zoology, and the Carnegie 
Museum, hundreds of other specimens were examined in the Museo 
Argentino de Ciencias Naturales. 
