632 FROVEEDINGS OF THE NATIONAL MUSEUM. vol. xxix. 



cell very iimch like the ambient vein of the Diptera. This condition 

 can be explained in another way b}^ assuming that the free part of R^, 

 the transverse part of M^, and the free part of M3 are all present, and 

 simply have migrated to the margin of the wing. The difficulty in 

 the way of this explanation is that the free part of K^ is always want- 

 ing in the females of these genera. In certain species of Phyllotoma 

 the free part of the second anal vein coincides with the second anal 

 furrow and the apex of the free part has atrophied. This subfamily 

 contains the genera, Galiroa^ PJiyllotoma^ and Erlociimpoides. 



THE SPECIALIZED TENTHREDINID^.. 



A group containing the greater part of the genera and species of 

 the family Tenth redinida\ They are differentiated from the gene- 

 ralized Tenthredinida' through the loss of the constriction near the 

 middle of the second anal cell. The subfamilies fall into several well- 

 marked lines of development. The Lycaotina? (tig. 55), Tenthredinina? 

 (tigs. 56-58), and Cimbicina? (tigs. 59-60) are the only members of this 

 series in which the free part of the second anal vein of the front wings is 

 preserved. The marked contraction of the third anal vein is repre- 

 sented as a slightl}' thickened emargination just before the free pai't of 

 the second anal vein, this is found only in the Lycaotinaj (tig, 55) and 

 Tenthredininaj. In the genera Macrophya (tig. 57) and Tenthredo (tig. 

 56) the presence or absence of a transverse vein between the second and 

 third anal veins is not of generic value. The Hoplocanjpiniv, L)inurina% 

 Cladiinaj, Monoctenina?, Nematint^?, Blennocampin*, Scolioneurin*, 

 and Fenusinjt have the anal cells either anastomosed at middle or with 

 the basal half of the third anal vein atrophied. In the Hylotomime, 

 Schizocerin*. and Perreyinse the second anal cell has been reduced by 

 the progressive coalesi-ence of the anastomosis to the base of the wing. 

 In the Lobocerina?, Pterygophorin*, and Perreyinse the anastomosis 

 has proceeded both ways, so that both the first and second anal cells 

 have been reduced. 



Lycdot'nfcV. — This subfamily contains the single genus Lycaota (tig. 

 65). Its location with the Tenthredinina> and Cimbicinaj is due to the 

 form of the anal cells, which anastomose at a single point at the usual 

 place for the location of the free part of the second anal vein. The 

 wings are broad and their apices are blunt. The medio-cubital cross- 

 vein arises from Sc + R+M, just before the origin of media. This 

 cross-vein and M.5+^ are parallel. In the hind wings the free part of 

 R^ is wanting and the first and second anal veins are coalesced for 

 nearly one-half the length of the veins. This subfamily is placed as 

 the most generalized member of the specialized Tenthredinida? because 

 of the form of the anal veins of the front wings and the position of 

 the medio-cubital cross-vein. 



