NO. 3556 PORTUNID CRABS—STEPHENSON AND REES 25 
last anterolateral tooth points in a similar direction, his species is 
here regarded (tentatively) as being a synonym of P. pseudoargentatus. 
Portunus gladiator (Alcock) 
Neptunus (Amphitrite) gladiator Alcock, 1899, pp. 35-36.—Shen, 1937, pp. 
101-108, figs. 2a-c. 
Monomia gladiator Barnard, 1950, p. 156. 
Portunus gladiator Crosnier, 1962, pp. 51-54, figs. 72, 76, 78, 82-83, pl. 3 (fig. 2). 
?Portunus gladiator Fabricius, 1798, p. 368. 
Not Portunus gladiator Stephenson and Campbell, 1959, pp. 110-111, figs. 2J, 
3J, pls. 3 (fig. 2), 41, 5J. 
Materiau.—Africa: Natal, from W. F. H. Rosenberg, 1 male 
(70 mm.), 1 female (66 mm.). 
Portunus pseudoargentatus Stephenson 
Portunus (Amphitrite) gladiator De Haan, 1835, p. 39, pl. 1 (fig. 5). 
Neptunus (Amphitrite) gladiator Sakai, 1939, pp. 390-391, text-fig. 5a, pl. 47 
fig. 3). 
a Sa iadnoke Stephenson, 196la, pp. 109-111, figs. 2A, 3F, pls. 
2 (fig. 4), 4F, 5D. 
?Portunus gladiator Fabricius, 1798, p. 368. 
MarterraAL.—Philippines: Lembeh Strait, Celebes, June 14, 1929, 
Herre collection, 1 male (25 mm.). 
Japan: Fukura, Awaji, Y. Hirase, 1 female (28 mm.); Shimizu, 
Suruga, shore, Oct. 14, 1906, Alb., 1 male (20 mm.). 
Remarxks.—This species differs in the following details from P. 
gladiator, as described and figured by Crosnier (1962) (the comparison 
has been facilitated by Crosnier’s kindness in forwarding one of his 
named specimens) : 
a. A more strongly embossed carapace with coarser granulation. 
In particular the cardiac, lateral postcardiacs, posterolaterals, and 
two mesobranchials are larger and less diffuse than in P. gladiator. 
b. Form of male abdomen. The ultimate segment relatively 
longer and without concave lateral borders, penultimate segment not 
swollen in distal third. 
c. Relatively narrower fifth merus. In P. pseudoargentatus 
breadth is about 0.75 times length, in P. gladiator about 0.9 times. 
d. Differently armed anterodistal border of fifth merus. In P. 
pseudoargentatus it is finely serrated, in P. gladiator it bears small 
but distinct teeth. 
e. The last anterolateral tooth is not directed slightly posteriorly, 
as in P. gladiator. 
Portunus granulatus (H. Milne Edwards) 
Figure 5 
Lupea granulata H. Milne Edwards, 1834, p. 454. 
Neptunus (Achelous) granulatus (H. Milne Edwards).—Alcock, 1899, pp. 45-46.— 
Sakai, 1939, p. 397, pl. 81 (fig. 3). 
