NO. 3656 PORTUNID CRABS—-STEPHENSON AND REES 29 
related species are: P. macrophthalmus Rathbun 1906, P. longi- 
spinosus bidens Laurie 1906, P. emarginatus Stephenson and Camp- 
bell, 1959, and P. tenuicaudatus Stephenson, 1961a. 
Disregarding synonymy problems, most of the undamaged males in 
the present collection could be referred to one or another of the above 
species without great difficulty. Problems arose over two dissimilar 
sized males (10 mm. and 20 mm. carapace breadths) from the same 
collection (Palau, Sta. 124-1183). Although undoubtedly belonging 
to the same species, they differed in certain features which have been 
relied upon heavily in past descriptions, viz the length and acuteness 
of the lateral frontal teeth, and the proportions and shapes of the 
ultimate and penultimate segments of the male abdomen. In the 
smaller specimen the frontal teeth are low and rounded (cf. P. emar- 
ginatus); the male abdomen is short and broad, with only slightly 
concave outer borders of the penultimate segment and with an 
ultimate segment as long as broad (cf. P. longispinosus Stephenson 
and Campbell). In the larger specimen the frontal teeth, while low 
and rounded, project farther (as in P. longispinosus Stephenson and 
Campbell), and the male abdomen, although moderately long and with 
distinctly concave borders to the penultimate segment (as in P. 
emarginatus), has the ultimate segment slightly longer than broad. 
P. longispinosus Stephenson and Campbell and P. emarginatus show 
many points of similarity, including an absence of tuberculate eleva- 
tions upon the granulated areas of the carapace and quite similar 
pleopods. Tentatively they are treated as synonyms, and specimens 
are listed under P. emarginatus. 
While, as indicated, frontal teeth and male abdomens may have 
less diagnostic importance than previously thought, they still have 
some value. Itis believed, however, that the details of ornamentation 
of the dorsal surface of the carapace offer the best opportunities for 
separating the species, although here again there is variation. 
In the mesobranchial areas of the carapace both of the present 
specimens of P. iranjae bear elevated tubercular projections that were 
not present in the specimens submitted by Crosnier to the senior 
author. Stephenson’s comments are quoted in Crosnier’s description. 
Recently Described Species in P. longispinosus Complex 
16 Upper surface of palm of chelipeds with a single distal spine. 
P. longispinosus bidens (Laurie) 1906 
Upper surface of palm of cheliped with two distal spines or with a spine 
carol G2} yopuatiitoraam EINE 4 5 5 6 GO oO-d Ob oO 6 On OO OOo 2 
2. (1) Outer frontal teeth low and rounded (granular eminences of carapace 
without tubercles). 
P. emarginatus Stephenson and Campbell, 1959 
Outer frontal teeth triangular, projecting, and typically sharp. . . 3 
