38 PROCEEDINGS OF THE NATIONAL MUSEUM VOL. 120 
(partim).—de Man, 1887, pp. 70-73 (under WN. andersoni).—Henderson, 
1893, p. 869 (under N. andersoni). 
Portunus rugosus (A. Milne Edwards).—Guinot, 1957, p. 480 (under P. mariez).— 
Stephenson, 1961la, pp. 111-114, figs. 2B, 3G, pls. 3 (fig. 1), 4G, 5E. 
InrRopuUcTION.—Original sorting of specimens gave seven different 
forms resembling P. rugosus. Differences between them concerned 
shape of frontal teeth, length of last anterolateral tooth, acuteness or 
otherwise of posterior-posterolateral junction of the carapace, oc- 
currence of a paried or single metagastric granular elevation of the 
carapace, curvature and spinulation of the pleopod, and relative 
length of penultimate segment of male abdomen. Later it was 
shown that some of these features varied with size of specimens 
within a single collection (e.g., length of last anterolateral tooth), 
while others varied between adjacent collections (e.g., acuteness or 
roundedness of frontal teeth). ‘To separate most of the remaining 
POST - ORBITAL POST FRONTAL 
MED. GASTRIC 
PROTOGASTRIC 
MESOGASTRIC 
ANT. ANTEROLAT, 
POST. ANTEROLAT. 
EPIBRANCHIAL 
ANT, MESOBRANCHIAL 
METAGASTRIC 
CENT. MESOBRANCHIAL 
CARDIAC 
POST. MESOBRANCHIAL 
LAT. POSTCARDIAC 
POSTEROLATERAL 
MED. POSTCARDIAC 
Ficure 8.—Portunus rugosus (A. Milne Edwards): Diagram of elevated granular areas of 
carapace. 
forms involved reliance upon a single character (e.g., paired or single 
metagastric elevation), and it was concluded that two forms, here 
designated I and II, were present. 
These were compared with the Western Australian material upon 
which Stephenson’s (1961a) redescription of the species is based. 
This material belongs to form I. 
The features common to all variants are listed below, followed by 
accounts of the variation within forms I and II and of the distinctions 
between them. ‘These are followed by a brief report upon Milne 
Edwards’ material. 
Marertau.—Form I: Philippines: Jolo, ship’s side, Feb. 8, 1908, 
Alb., 1 female (dried); Jolo anchorage, electric light, Feb. 13, 1908, 
Alb., 1 male; Sta. 5156, Tinakta I., Sulu Archipelago, Tawitawi 
Group, 5°12’50”N., 119°55’55”E., Feb. 21, 1908, Alb., 1 male; 
Sta. 5157, Tinakta I., Sulu Archipelago, Tawitawi Group, 5°21’307N., 
119°55’50”E., 18 fm., fine sand, Feb. 21, 1908, Alb., 5 males, 3 
