NO. 3556 PORTUNID CRABS—STEPHENSON AND REES 53 
Sta. 5159, Tinakta I., Sulu Archipelago, Tawitawi Group, 5°11’50” 
N., 119°54’E ., Feb. 21, 1908, Alb., 1 male; Varadero Bay, Mindanao, 
10:30-11:30, electric light, July 22, 1908, Alb., 1 female; Sta. 5442, 
west coast of Luzon, South Fernando Point Light, 16°30’36’N.., 
120°11’06”E., 45 fm., coarse sand, May 11, 1909, Alb., 7 males, 1 
female, 2 ovig. females; Sta. 5478, Tacbuc Point, (Leyte), 10°46’24” 
N., 125°16’30’E., July 29, 1909, Alb., 1 female; Sta. 5642, Buton 
Strait, 4°31’40"S., 122°49’42”K., 37 fm., grey mud, Dec. 14, 1909, 
Alb., 1 male. 
Palau: Sta. 125-1187 Palau I., about 134 mi. NE. of Ngabadongu, 
7°17'36"N., 134°21/42”E., 17 fm., Aug. 24, 1955, GVF., 1 female. 
MeAsurREMENTS.—Males, 18-49 mm.; females 15-30 mm.; ovig. 
females, 21-26 mm. 
Remarxs.—Originally it was suspected that two different species 
were present, some being more strongly embossed and_ possessing 
spiniform tubercles on the cardiac and lateral postcardiac regions of 
the carapace. Later intergrades were found. The following, the 
only adequate description in English, amplifies Crosnier’s (1962) 
French description. 
Description.—Front: Projecting, four round, approximately equal 
lobes, medians the more prominent. 
Anterolateral teeth: Irregularly sized and spaced, first and second 
almost confluent, fourth and sixth small and depressed, third and fifth 
with enlarged granular bases, ninth the longest. 
Carapace: Moderately broad (B/L.=1.6-1.7). Regions varying 
from moderately to very strongly embossed, and carrying beaded 
granules or occasionally tubercles. Strongly depressed posterior to 
protogastric and mesogastric areas. Main granular patches as fol- 
lows: Protogastrics subdivided into two; mesogastrics, typically two 
oblique patches; metagastrics, transverse ridge with anterior prolonga- 
tion; cardiacs either an elevated patch, or pair of patches, or pair of 
elevated granular tubercles; lateral postcardiacs elevated and some- 
times bearing tubercles; median postcardiac present; anterolaterals 
diffuse; epibranchials with marked change in curvature near middle 
and interrupted on cervical groove to form discrete patch; three meso- 
branchials; diffuse posterolateral. 
Chelipeds: Elongate, right slightly the stouter. Upper surfaces cov- 
ered with beaded granules and lower surfaces, particularly of hands, 
bearing squamiform markings. Anterior border of arm bearing two 
spines typically followed by a tubercle, occasionally by a third spine. 
Posterodistal border with single spine. (Crosnier describes two teeth, 
the second ‘“méritant 4 peine le nom de dent.) Wrist normal. 
Upper surface of hand bearing three granular carinae of which inner- 
most terminates in distal spine or spiniform tubercle. Conspicuous 
