NO. 3558 MALLOPHAGA—ELBEL 43 
USNM skins from Tarussan Bay, West Sumatra, 1904-1905, collected 
by W. L. Abbott, REE; from Buceros rhinoceros borneoensis Schlegel 
and Miiller, 1840: 1 male and 6 females from Serabang Bay, Sarawak, 
Borneo, Jan. 11, 1958, BMNH 1958-737; 4 females from Borneo, 
Meinertzhagen 10890, BMNH; 2 males and 5 females from CNHM 
skin from Sapagayo Forest Reservation, Sandakan, North Borneo, July 
27, 1950, collected by R. F. Inger and D. D. Davis, REE; from 
Buceros bicornis homrai Hodgson, 1832: 7 males and 5 females from 
Ban Khlua Klang, Prachuap Khiri Khan, Thailand, December 1952, 
collected by Robert E. Elbel, and H. G. Deignan, USNM; 11 males 
and 8 females from Khlong Khlung, Kamphaeng Phet, Thailand, 
Apr. 7, 1953, collected by Robert E. Elbel and H. G. Deignan, USNM; 
1 male and 3 females from Ban Muang Khai, Tha Li, Loei, Thailand, 
Dec. 7, 1953, collected by Robert E. Elbel, USNM; 2 males and 1 
female from Banghin, Kapoe, Ranong, Thailand, Feb. 6, 1963, 
collected by Wichit Suwan Laong, USNM; from Rhinoplax vigil 
(J. R. Forster, 1781): 1 female from Borneo, Meinertzhagen 10888, 
BMNH. 
Drawings were made of a male and a female from Buceros bicornis 
homrai collected in Khlong Khlung, Thailand. Specimens in USNM. 
The productus Species-Group 
Species similar in shape to Bucerophagus africanus (figs. 68, 69). 
Differing from the forcipatus species-group in the following combina- 
tion of characters: Head width 1% to 1% times that of length; metano- 
tum with 3-5 short setae on each lateral margin and with 6-8 setae 
on anterior margin; metasternal plate with more than 20 setae; 
venter of third femora and posterolateral margins of abdominal 
sternites IV and V each with thick brushes of normal setae; each lateral 
margin of abdominal tergites JI-VIII with 1-5 short setae between 
the spiracle and postspiracular seta; females with more abdominal 
sternal setae than males; both sexes having abdominal sternite I 
with more than 20 total setae and abdominal sternite IL with more 
than 60 total setae; male terminal abdominal sternites VIII and IX 
neither projecting posteriorly as rounded lobe nor with division but 
with complete division from abdominal sternite VII; male genitalia 
with parameres straight and not split posteriorly; female terminal 
abdominal tergite with more than 24 setae on posterior margin; 
female abdominal sternite VIII with more than 28 setae on posterior 
margin. 
Hosts: Species of the productus species-group have been found 
only on the genus Bucorvus of the avian family Bucerotidae. 
