NO. 3567 LABIDOCERA JOLLAE GROUP—FLEMINGER 37 
ranges. Adults of this group show two characteristics which set them 
apart from the remainder of the genus, namely: (1) antennules with 
segments 8 and 9 partially fused; and (2) mandibular gnathobase with 
basal teeth 1 and 2 usually bicuspidate. The jollae group is further 
distinguished from their closest relatives, the mirabilis group 
(Fleminger and Tan, 1966) in the retention of head-hooks (except in 
adult diandra). 
A large number of morphological features are shared exclusively 
by the mirabilis and jollae groups. This and their endemism in the 
Americas constitute a firm basis for combining them in the wilsoni 
superspecies as proposed by Fleminger and Tan, (1966). This super- 
species is characterized by (1) short rostral prongs, which have a 
swollen proximal portion and are separated in frontal view by a 
triangular notch (fig. 10); (2) separation of TIV and TV; (3) two- 
segmented, asymmetrical urosome and usually asymmetrical caudal 
rami in the female; (4) male left leg 5 usually short, terminal segment 
padlike, densely pilose and bearing three minute spines or setae or 
none at all; (5) male lacking specialized outgrowths on right corner 
of TV; and (6) male with asymmetrical exopodal spines on one or 
more pairs of swimming legs. Also noteworthy is the strong overall 
similarity of immature stages, especially in such critical features as the 
fifth legs and rostrum and the crimson bodies in the thorax. 
Other head-hooked species differ from the jol/ae group in having 
(based on various sources including unpublished observations of the 
author) (1) relatively long, slender rostral prongs; (2) nongeniculated 
antennule segmentation numbering 24(24—25 fused) or, if 23, with 
6-7 and 24-25 fused; (3) female with three-segmented urosome; 
(4) male left leg 5 more than one half the length of right leg 5, terminal 
segment of former with several robust spiniform or setiform processes, 
conspicuous and often incompletely fused to segment; (5) male TV 
usually asymmetrical, right posterior corner being variously modi- 
fied; and (6) male right antennule with segment 17 bearing modified 
seta anteriad, more or less ridgelike. 
The wilsont superspecies shows morphological affinities with another 
assemblage of predominantly Indo-Pacific species of which L. detrun- 
cata is characteristic. This resemblance is restricted to a few struc- 
tures modified by secondary sexual development and lacks the support 
of similarities in rostrum configuration and Al segmentation. In 
reviewing the genus, division of the species among groups and super- 
species has proven most helpful and provisional monophyletic aggre- 
gations are given below. 
Superspecies wilsoni (coastal, temperate to subtropical). 
jollae group: L. jollae Esterly, diandra, new species, kolpos, new species (west 
coast, North America). 
