Ceylon and the. Philippine Islands. 97 



known at present, unique among the Oligochfeta. There are 

 two pairs of testes in the 8th and 10th segments respectively ; 

 each of these opens on to the exterior by a separate vas deferens 

 furnished at its termination with a prostate gland ; the ante- 

 rior pair of apertures are placed in the 7th segment in front 

 of the testes belonging to them, the latter between the lOtli 

 and 11th segments behind their testes. This latter pair of 

 apertures correspond exactly to those which I have found in 

 M. Barwelli. The anterior pair of testes together with the 

 accessory structures were entirely absent ; but in this segment 

 were a pair of copulatory pouches (fig. 5) opening on to the 

 exterior in front of the outermost pair of setse. The copula- 

 tory pouches are remarkable from the fact that the pouch 

 itself is a small spherical vesicle communicating with the ex- 

 terior by a long, slender, variously coiled and contorted duct, 

 which, together with the pouch, is closely applied to the 

 mesentery dividing the segment from the one in front. The 

 length of the duct is remarkable and recalls the copulatory 

 pouch of certain Limicolous Oligochseta (e. g. Anachceta Eiseni, 

 Vejdovsky, loc. cit. pi. vii. fig. 22). The presence of a pair 

 of copulatory pouches, instead of an anterior pair of testes 

 and vasa deferentia, brings tiie structure of the genital appa- 

 ratus in this species much nearer to the usual condition met 

 with in earthworms. M. Perrier's account of the male genital 

 apparatus of M. Deshayesi, which is illustrated, is too circum- 

 stantial to admit of any doubts of its accuracy, though there is 

 evidently some resemblance between the vas deferens (" entor- 

 tillde corame serait un Gordius ") of M. Deshayesi and the 

 convoluted duct of the copulatory pouch in the present species. 

 I feel uncertain therefore whether there is a real difference 

 between the two species in the structure of the male genital 

 organs, or whether the anterior testes and their ducts may not, 

 as M. Perrier suggests, be developed at different times ; at the 

 same time it seems hardly likely that the vasa deferentia and 

 prostates would disappear with the testes, and I am certain 

 that a second pair of these structures did not exist in the speci- 

 mens of M. Barioelli which I have examined. Moreover, a 

 comparison of M. Perrier's figures of the posterior testes with 

 their ducts and accessory structures * with my own (fig. 4) 

 will show that there is sufficient difference to separate the two 

 forms, at least specifically, without reference to the presence 

 or absence in M. Barwelli of the anterior testes, 



* Loc. cit. pi, iv. figs. 79 and 81. 



