﻿PROCEEDINGS 
  OF 
  THE 
  UNITED 
  STATES 
  NATIONAL 
  MUSEUM 
  

  

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  SMITHSONIAN 
  INSTITUTION 
  

   U. 
  S. 
  NATIONAL 
  MUSEUM 
  

  

  Vol.103 
  Washington 
  : 
  1953 
  No. 
  3318 
  

  

  SCARABAEID 
  BEETLES 
  OF 
  THE 
  GENUS 
  BRADYCINETULUS 
  

   AND 
  CLOSELY 
  RELATED 
  GENERA 
  IN 
  THE 
  UNITED 
  STATES 
  

  

  By 
  O. 
  L. 
  Cartwright 
  

  

  The 
  group 
  of 
  beetles 
  reviewed 
  in 
  this 
  paper 
  have 
  long 
  been 
  known 
  

   under 
  the 
  name 
  Bolboceras. 
  The 
  species 
  found 
  in 
  the 
  United 
  States 
  

   were 
  last 
  revised 
  by 
  Charles 
  Schaeffer, 
  1906 
  (Trans. 
  Amer. 
  Ent. 
  Soc, 
  

   vol. 
  32, 
  pp. 
  249-254). 
  At 
  that 
  time 
  five 
  species 
  were 
  placed 
  in 
  Horn's 
  

   genus 
  Bradycinetus, 
  later 
  incorrectly 
  synonymized 
  under 
  Bolboceras 
  

   (see 
  synonymy 
  under 
  Bradycinetulus) 
  , 
  and 
  three 
  were 
  assigned 
  to 
  

   Bolboceras. 
  Of 
  these 
  three, 
  Scarabaeus 
  lazarus 
  Fabricius 
  had 
  been 
  

   placed 
  in 
  Eucanthus 
  by 
  Westwood, 
  1848 
  (Proc. 
  Linn. 
  Soc. 
  London, 
  

   vol. 
  1, 
  p. 
  387), 
  and 
  Bradycinetus 
  hornii 
  Rivers 
  and 
  B. 
  minor 
  Linell 
  

   were 
  later 
  removed 
  to 
  Kolbeus 
  by 
  Chapin 
  (see 
  synonymy 
  under 
  

   Bolbelasmus) 
  . 
  

  

  With 
  the 
  exception 
  of 
  the 
  species 
  Eucanthus 
  and 
  Bolbocerosoma, 
  here 
  

   considered 
  correctly 
  placed 
  in 
  recent 
  usage, 
  all 
  of 
  the 
  United 
  States 
  

   species 
  listed 
  by 
  Schaeffer 
  have 
  been 
  reexamined 
  in 
  this 
  study 
  and 
  

   reassigned 
  to 
  genera 
  as 
  required 
  by 
  the 
  literature 
  on 
  the 
  group. 
  Also 
  

   included 
  in 
  this 
  paper 
  are 
  the 
  one 
  species 
  described 
  siiice 
  Schaeffer's 
  

   time, 
  Bolboceras 
  angulus 
  Robinson; 
  a 
  Mexican 
  species 
  discovered 
  in 
  the 
  

   United 
  States; 
  another 
  which 
  may 
  eventually 
  be 
  found 
  inside 
  our 
  

   borders; 
  and 
  four 
  new 
  United 
  States 
  species. 
  Two 
  new 
  genera 
  are 
  

   proposed. 
  A 
  key, 
  based 
  in 
  part 
  on 
  that 
  by 
  Boucomont, 
  is 
  presented 
  

   for 
  the 
  genera 
  of 
  the 
  Bolboceratini 
  found 
  in 
  the 
  United 
  States. 
  The 
  

   genera 
  Kolbeus, 
  Bolbelasmus, 
  Bradycinetulus, 
  and 
  the 
  two 
  new 
  genera 
  

   Bolbocerastes 
  and 
  Bolborhombus 
  are 
  discussed, 
  synonymy 
  listed, 
  and 
  a 
  

   key 
  to 
  the 
  species 
  given. 
  Pertinent 
  references, 
  the 
  type 
  locality 
  and 
  

   location 
  of 
  the 
  type, 
  and 
  notes 
  on 
  distribution 
  follow 
  for 
  each 
  species. 
  

   Distribution 
  is 
  also 
  shown 
  on 
  the 
  maps, 
  figures 
  15 
  and 
  16. 
  The 
  genus 
  

  

  232990—53 
  1 
  95 
  

  

  