﻿166 
  PROCEEDINGS 
  OF 
  THE 
  NATIONAL 
  MUSEUM 
  vol. 
  io3 
  

  

  It 
  is 
  lined 
  with 
  a 
  tall, 
  glandular 
  epithelium, 
  the 
  cells 
  of 
  which 
  project 
  

   into 
  the 
  cavity 
  in 
  a 
  villuslike 
  fashion. 
  Below 
  the 
  epithelium 
  there 
  

   are 
  two 
  layers 
  of 
  muscle 
  fibers, 
  one 
  circular 
  and 
  the 
  other 
  longitudinal. 
  

  

  The 
  penis 
  consists 
  of 
  a 
  spherical, 
  muscular 
  bulb 
  embedded 
  in 
  the 
  

   mesenchyme, 
  and 
  a 
  moderately 
  large 
  papilla 
  projecting 
  into 
  the 
  male 
  

   atrium. 
  The 
  bulb 
  is 
  pierced 
  by 
  numerous 
  gland 
  ducts 
  which 
  open 
  

   into 
  the 
  lumen 
  of 
  both 
  the 
  bulb 
  and 
  the 
  papilla. 
  The 
  shape 
  of 
  the 
  

   papilla 
  is 
  subject 
  to 
  great 
  variation, 
  due 
  apparently 
  to 
  the 
  state 
  of 
  

   contraction 
  of 
  the 
  organ. 
  It 
  may 
  be 
  twisted 
  to 
  one 
  side 
  and 
  even 
  

   partly 
  inverted 
  into 
  the 
  lumen 
  of 
  the 
  penis 
  (similar 
  to 
  the 
  pseudo- 
  

   flagellum 
  of 
  various 
  dendrocoelids) 
  . 
  The 
  shape 
  shown 
  in 
  figure 
  21 
  

   appears 
  to 
  be 
  that 
  of 
  the 
  organ 
  at 
  rest. 
  The 
  outer 
  wall 
  of 
  the 
  papilla 
  

   is 
  covered 
  with 
  a 
  tall 
  to 
  cubical 
  epithelium 
  similar 
  to 
  that 
  lining 
  the 
  

   atrium. 
  Below 
  the 
  epithelium 
  there 
  is 
  a 
  layer 
  of 
  circular 
  muscle 
  

   fibers 
  followed 
  by 
  another 
  of 
  longitudinal 
  fibers. 
  The 
  shape 
  of 
  the 
  

   penis 
  lumen 
  {Ip) 
  is 
  as 
  changeable 
  as 
  that 
  of 
  the 
  papilla. 
  Typically, 
  

   it 
  appears 
  to 
  be 
  wider 
  in 
  the 
  bulb 
  than 
  it 
  is 
  in 
  the 
  papilla, 
  though 
  

   there 
  is 
  no 
  distinct 
  ejaculatory 
  duct 
  differentiated. 
  The 
  lumen 
  opens 
  

   ventrally 
  to 
  the 
  tip 
  of 
  the 
  papUla. 
  The 
  two 
  vasa 
  deferentia 
  {vd) 
  

   penetrate 
  the 
  penis 
  bulb 
  from 
  both 
  sides 
  and 
  empty 
  into 
  the 
  penis 
  

   lumen 
  separately, 
  but 
  not 
  far 
  apart. 
  

  

  The 
  two 
  oviducts 
  converge 
  at 
  the 
  level 
  of 
  the 
  copulatory 
  apparatus, 
  

   the 
  left 
  one 
  passing 
  between 
  the 
  bursa 
  duct 
  and 
  the 
  male 
  atrium, 
  and 
  

   unite 
  at 
  a 
  point 
  dorsally 
  to 
  the 
  atrium. 
  The 
  rather 
  long 
  common 
  

   oviduct 
  {ode) 
  curves 
  ventrally 
  and 
  opens 
  into 
  the 
  posterior 
  part 
  of 
  the 
  

   male 
  atrium. 
  The 
  terminal 
  sections 
  of 
  the 
  paired 
  oviducts 
  and 
  the 
  

   greater 
  part 
  of 
  the 
  common 
  oviduct 
  receive 
  the 
  outlets 
  of 
  numerous 
  

   eosinophilic 
  glands, 
  the 
  cell 
  bodies 
  of 
  which 
  are 
  scattered 
  in 
  the 
  sur- 
  

   rounding 
  mesenchyme, 
  particularly 
  dorsally 
  to 
  the 
  atrium. 
  

  

  The 
  copulatory 
  bursa 
  (6) 
  is 
  of 
  moderate 
  to 
  large 
  size 
  and 
  is 
  ir- 
  

   regularly 
  lobed. 
  The 
  bursa 
  duct 
  or 
  stalk 
  {hd) 
  is 
  wide, 
  runs 
  posteriorly 
  

   to 
  the 
  left 
  of 
  the 
  midline, 
  and 
  curves 
  ventrally 
  to 
  reach 
  the 
  genital 
  

   aperture. 
  It 
  is 
  lined 
  with 
  a 
  tall, 
  glandular 
  epithelium 
  and 
  surrounded 
  

   with 
  a 
  strong 
  muscular 
  coat 
  consisting 
  of 
  intermingled 
  circular 
  and 
  

   longitudinal 
  fibers. 
  

  

  Taxonomic 
  position. 
  — 
  The 
  genus 
  Phagocata 
  Leidy 
  (Fonticola 
  

   Komarek) 
  in 
  its 
  present 
  extent 
  (cf. 
  Hyman, 
  1937, 
  pp. 
  300-302) 
  has 
  

   representatives 
  in 
  Europe, 
  Asia, 
  and 
  North 
  America. 
  The 
  genus 
  is 
  

   not 
  quite 
  homogeneous 
  and 
  will 
  probably, 
  in 
  due 
  time, 
  be 
  subdivided 
  

   into 
  several 
  genera 
  (cf. 
  Beauchamp, 
  1939). 
  For 
  the 
  purpose 
  of 
  com- 
  

   parison, 
  we 
  may 
  consider 
  here 
  only 
  those 
  species 
  of 
  the 
  genus 
  that 
  

   lack 
  pigment. 
  Though 
  the 
  presence, 
  or 
  the 
  lack, 
  of 
  pigment 
  is 
  a 
  

   character 
  of 
  subordinate 
  taxonomic 
  value, 
  it 
  may 
  nevertheless 
  serve 
  

   well 
  as 
  a 
  character 
  of 
  specific 
  rank. 
  The 
  Alaskan 
  form 
  differs 
  from 
  

  

  