﻿CANYON 
  FERRY 
  FOSSIL 
  VERTEBRATES 
  — 
  ^WHITE 
  407 
  

  

  the 
  same 
  species. 
  The 
  two 
  specimens 
  are 
  from 
  the 
  same 
  horizon 
  and 
  

   locaHty 
  but 
  the 
  difference 
  in 
  amount 
  of 
  wear 
  excludes 
  them 
  from 
  

   belonging 
  to 
  the 
  same 
  individual. 
  

  

  Discussion. 
  — 
  Although 
  this 
  specimen 
  adds 
  many 
  new 
  data 
  to 
  our 
  

   knowledge 
  of 
  this 
  group 
  of 
  insectivores, 
  the 
  affinities 
  of 
  the 
  family 
  are 
  

   still 
  problematical. 
  A 
  number 
  of 
  the 
  characters 
  are 
  shared 
  by 
  both 
  

   the 
  Soricoidea 
  and 
  the 
  Microchiroptera. 
  Some 
  characters 
  are 
  shared 
  

   by 
  some 
  members 
  of 
  both 
  groups 
  but 
  not 
  by 
  all 
  members 
  of 
  either. 
  

   Briefly, 
  these 
  characters 
  are 
  the 
  fused 
  skull 
  elements 
  and 
  the 
  char- 
  

   acter 
  of 
  the 
  orbits. 
  This 
  is 
  also 
  true 
  for 
  the 
  general 
  characters 
  of 
  the 
  

   cheek 
  teeth. 
  In 
  some 
  members 
  of 
  both 
  groups 
  the 
  posterior 
  border 
  

   is 
  deeply 
  emarginate 
  between 
  the 
  hypocone 
  and 
  metastyle. 
  The 
  

   parastylar 
  and 
  metastylar 
  crests 
  resemble 
  both 
  groups 
  in 
  their 
  general 
  

   features 
  but 
  in 
  their 
  details 
  resemble 
  the 
  bats 
  more 
  than 
  the 
  shrews. 
  

   The 
  very 
  high 
  cusps 
  of 
  the 
  cheek 
  teeth, 
  especially 
  the 
  hypocone, 
  

   appear 
  to 
  be 
  a 
  unique 
  feature 
  of 
  this 
  specimen. 
  The 
  large 
  canine 
  

   excludes 
  this 
  specimen 
  from 
  the 
  shrews 
  but 
  not 
  from 
  the 
  moles 
  (Talpinae) 
  

   and 
  bats. 
  The 
  elongate 
  pit 
  in 
  front 
  of 
  the 
  orbit 
  is 
  a 
  characteristic 
  of 
  

   most 
  shrews 
  and 
  many 
  of 
  the 
  moles, 
  but 
  not 
  of 
  the 
  bats. 
  A 
  slender 
  

   zygoma 
  is 
  present 
  in 
  moles 
  and 
  most 
  bats 
  but 
  is 
  absent 
  in 
  the 
  shrews. 
  

   The 
  rostrum 
  is 
  broken 
  away 
  and 
  the 
  anterior 
  termination 
  of 
  the 
  face 
  

   cannot 
  be 
  determined. 
  

  

  In 
  summary, 
  the 
  characters 
  of 
  this 
  specimen 
  exclude 
  it 
  from 
  the 
  

   Soricidae 
  but 
  do 
  not 
  definitely 
  affiliate 
  it 
  with 
  any 
  other 
  group. 
  The 
  

   molelike 
  facial 
  features 
  certainly 
  suggest 
  that 
  it 
  could 
  belong 
  to 
  the 
  

   talpid 
  humerus 
  from 
  the 
  same 
  locality. 
  Parascaptor 
  and 
  Scapato- 
  

   chirus 
  have 
  a 
  large 
  canine 
  and 
  reduced 
  premolars 
  as 
  does 
  this 
  specimen. 
  

   However, 
  the 
  cheek 
  teeth 
  of 
  this 
  specimen 
  are 
  of 
  a 
  type 
  unknown 
  in 
  

   any 
  living 
  moles 
  and 
  very 
  suggestive 
  of 
  the 
  bats. 
  Measurements 
  of 
  

   teeth 
  (in 
  millimeters) 
  : 
  

  

  USNM 
  18870 
  

  

  C-M3 
  11. 
  6 
  

  

  P3-P* 
  3. 
  6 
  

  

  M»-M3 
  5. 
  5 
  

  

  Length 
  Width 
  

  

  P* 
  2.2 
  1.8 
  

  

  Ml 
  2.3 
  2.5 
  

  

  M2 
  1.8 
  2.7 
  

  

  M? 
  1.1 
  2.2 
  

  

  USNM 
  18871 
  

  

  Width 
  Height 
  

  

  Length 
  Trigonid 
  Heel 
  Trigonid 
  Heel 
  

  

  Mi 
  1.8 
  1.8 
  1.2 
  3.1 
  1.3 
  

  

  M3 
  1.8 
  1.5 
  0.8 
  2.1 
  1.0 
  

  

  Depth 
  of 
  jaw 
  between 
  M2 
  and 
  M3 
  2. 
  3 
  

  

  Depth 
  of 
  jaw 
  below 
  masseteric 
  fossa 
  1. 
  2 
  

  

  