﻿ANATOMY 
  AND 
  RELATIONSHIP 
  OF 
  GLOSSY 
  CUCKOOS 
  — 
  BERGER 
  587 
  

  

  Osteology 
  

  

  All 
  four 
  species 
  have 
  14 
  cervical 
  vertebrae. 
  There 
  are 
  two 
  pairs 
  

   of 
  cervicodorsal 
  ribs 
  and, 
  usually, 
  only 
  the 
  last 
  such 
  rib 
  possesses 
  an 
  

   uncinate 
  process, 
  though 
  one 
  specimen 
  of 
  Chrysococcyx 
  cupreus 
  has 
  

   uncinate 
  processes 
  on 
  both 
  left 
  cervicodorsal 
  ribs, 
  one 
  specimen 
  of 
  

   Lam'promorpha 
  caprius 
  possesses 
  these 
  processes 
  on 
  both 
  right 
  ribs, 
  

   and 
  an 
  alcoholic 
  specimen 
  of 
  Chalcites 
  lucidus 
  has 
  uncinate 
  processes 
  

   on 
  all 
  four 
  ribs. 
  The 
  atlas 
  is 
  perforated 
  by 
  the 
  odontoid 
  process, 
  ex- 
  

   cept 
  in 
  a 
  strongly 
  bleached 
  skeleton 
  of 
  Lampromorpha 
  klaas, 
  where 
  

   the 
  atlas 
  appears 
  to 
  be 
  notched. 
  

  

  With 
  one 
  exception, 
  each 
  specimen 
  has 
  four 
  dorsal 
  vertebrae, 
  four 
  

   dorsal 
  ribs, 
  and 
  one 
  thoracic 
  rib. 
  In 
  each 
  case, 
  the 
  thoracic 
  rib 
  articu- 
  

   lates 
  with 
  the 
  synsacrum 
  dorsally 
  and 
  fuses 
  with 
  the 
  last 
  dorsal 
  rib 
  

   ventrally. 
  One 
  alcoholic 
  specimen 
  of 
  Chrysococcyx 
  cupreus^ 
  however, 
  

   has 
  five 
  dorsal 
  vertebrae 
  and 
  five 
  dorsal, 
  or 
  true, 
  ribs. 
  Of 
  the 
  latter, 
  

   four 
  articulate 
  with 
  the 
  sternum, 
  while 
  the 
  fifth 
  fuses 
  ventrally 
  with 
  

   the 
  fourth 
  rib 
  ; 
  there 
  is 
  no 
  thoracic 
  rib. 
  Of 
  ten 
  cuculine 
  genera 
  I 
  have 
  

   thus 
  far 
  investigated, 
  this 
  is 
  the 
  first 
  example 
  I 
  have 
  seen 
  of 
  intra- 
  

   specific 
  variation 
  in 
  the 
  number 
  of 
  dorsal 
  vertebrae. 
  The 
  number 
  

   of 
  ribs 
  articulating 
  directly 
  with 
  the 
  sternum, 
  on 
  the 
  other 
  hand, 
  is 
  

   far 
  more 
  variable. 
  Four, 
  apparently, 
  is 
  the 
  "normal" 
  number 
  of 
  

   sternal 
  ribs 
  in 
  these 
  species. 
  In 
  one 
  specimen 
  of 
  Chrysococcyx 
  cupreus 
  

   and 
  two 
  specimens 
  of 
  Lampromorpha 
  caprius^ 
  however, 
  only 
  three 
  of 
  

   the 
  four 
  dorsal 
  ribs 
  articulate 
  directly 
  with 
  the 
  sternum; 
  and 
  in 
  an 
  

   alcoholic 
  specimen 
  of 
  L. 
  hlaas 
  three 
  ribs 
  articulate 
  with 
  the 
  sternum 
  

   on 
  the 
  left 
  side, 
  whereas 
  four 
  do 
  so 
  on 
  the 
  right. 
  

  

  Apparently 
  in 
  younger 
  individuals 
  the 
  general 
  pattern 
  of 
  the 
  pos- 
  

   terior 
  emargination 
  of 
  the 
  sternum 
  is 
  double-notched 
  ; 
  it 
  does 
  not 
  seem 
  

   likely 
  that 
  the 
  bone 
  would 
  be 
  resorbed 
  to 
  form 
  fenestrae 
  in 
  older 
  

   individuals. 
  In 
  half 
  of 
  the 
  specimens, 
  the 
  sternum 
  is 
  single-notched 
  

   and 
  bilateral 
  fenestrae 
  are 
  present. 
  In 
  some 
  cases, 
  these 
  fenestrae 
  are 
  

   bounded 
  posteriorly 
  by 
  a 
  bony 
  bar 
  only 
  1 
  mm, 
  in 
  width. 
  

  

  The 
  external 
  nares 
  are 
  oval 
  to 
  rounded 
  and 
  their 
  borders 
  are 
  con- 
  

   spicuously 
  swollen. 
  On 
  the 
  floor 
  of 
  the 
  narial 
  cavity 
  is 
  a 
  well-de- 
  

   veloped 
  tubercle 
  which 
  has 
  a 
  small 
  bony 
  core; 
  the 
  latter 
  is 
  readily 
  

   discernible 
  in 
  the 
  skeleton. 
  

  

  A 
  pectineal 
  process 
  is 
  not 
  developed. 
  

  

  Wing 
  Myology 
  

  

  In 
  all 
  four 
  species 
  the 
  following 
  wing 
  muscles 
  are 
  absent: 
  pro- 
  

   scapulohumeralis 
  brevis, 
  abductor 
  indicis 
  brevis, 
  abductor 
  digiti 
  II, 
  

   and 
  the 
  biceps 
  slip 
  to 
  the 
  tendon 
  of 
  M. 
  tensor 
  patagii 
  longus. 
  Gadow's 
  

   pars 
  propatagialis 
  musculi 
  cucullaris 
  does 
  not 
  insert 
  on 
  the 
  tendon 
  of 
  

   M. 
  tensor 
  patagii 
  longus. 
  

  

  