ATLANTIC AND CARIBBEAN PYCNOGONIDA—HEDGPETH 219 
Anepiua pallida, might be collected which would clarify the status 
of Say’s genus. Since that time more than 30 species have been 
referred to Anoplodactylus, and Norman’s attempt to reinstate 
Anaphia was ignored by taxonomists (with the exception of Carpenter, 
1912) and has received but one recognition in ecological literature 
(Lebour, 1916). In her most recent paper Lebour (1945) returns to 
Anoplodactylus without comment. 
According to the collections studied for this review, Anoplodactylus 
lentus Wilson is the commonest species of pycnogonid on the coast of 
South Carolina, but Phoxichilidium femoratum (Rathke), the species 
to which Say’s description might also apply inasmuch as it was based 
on female specimens, does not occur south ef Long Island Sound (see 
fig. 26). Say’s types appear to be no longer in existence, which makes 
it impossible to settle the matter with absolute finality.21 Even if the 
types were still extant, however, more would be lost than gained by 
restoring Say’s genus. Such a procedure would cause more unneces- 
sary confusion in a group which has already had more than its share 
of such unhappy taxonomy. Therefore it is urged that Anoplodactylus 
Wilson be retained and that Anaphia Say be rejected from further 
consideration, and be relegated to the status of a nomen odlitum, since 
it has not been in general use for more than 30 years. It is understood 
that this modification in the International Rules may be considered 
by the next committee.” 
The principal character of this genus is the prominently developed 
neck. The oviger is usually 6-jointed (with a nonarticulated seg- 
mentation on the third joint in many species). Wilson considered 
the apparent absence of auxiliary claws a generic character and coined 
his name to describe that condition, but minute auxiliary claws are 
present in most of the species, including Anoplodactylus lentus. The 
femoral cement glands of the males are of several types: In A. lentus 
the opening is a long slit on the median dorsal surface; in A. peiiolatus 
it is a short, tubular projection; and in A. insignis it is an oval cribri- 
form aperture. Still another form, which seems to be transitional 
between the tubular and cribriform types, is found in A. quadratispino- 
sus. This is a curious transparent vesicle (fig. 32, 6), with a pore at 
the summit. 
The females of many species have peculiar processes or tubercles 
on the ventral surface of the proboscis. In some species these are 
simple knobs; in others they are elaborate lobed processes. Their 
function and anatomical significance are obscure; Calman (1923, 
p. 289) has pointed out that they cannot be considered embryonic 
21 T am indebted to Dr. Richard A. McLean, who kindly searched the Say collections at the Academy of 
Natural Sciences of Philadelphia in my behalf for Say’s types. How fortunate that Rafinesque described 
no pycnogonids! 
22 See Ernest Mayr, ‘‘Systematics and the Origin of Species,’ p. 17, 1942, 
