ON A NEW SUBFAMILY OF PHYLLOSTOME BATS. 
BY 
Harrison ALLEN, M. D. 
IT have concluded that Natalus should be removed from the Vesper- 
tilionide and should constitute a subfamily of the Phyllostomidz. The 
characters of the Nataline are as follows: The nose-leaf absent in the 
adult, but rudimental in the late stage of the embryo. It forms an 
ovate elevation placed with long axis between the nostrils and reaches 
from the lip-margin to a short distance above the upper edge of the 
muzzle. <A well-marked vertical line is seen in the third and fourth in- 
terdigital spaces, extending nearly their entire lengths. The vomer is 
produced and continuous with the crest of the basisphenoid bone; the 
sides of the mesopterygoid fossa are deeply concave. The ectoturbinal 
is a mere rudiment. Three premolars are present, both above and be- 
low, and six incisors below and four above, a wide interval separating 
the centrals. The cuboid bone projects plantarly; two phalanges are 
present in the third manal digit. The prosternum, first pair of ribs, 
and intercostal cartilages are co-ossified, much the same manner as in 
the remote Hipposideros. There is no terminal cartilage to the third 
manal digit. 
The claim here made that the Nataline are related to the Vampyri- 
group of the Phyllostomide rest& on the following statements: A rudi- 
ment of a nose-leaf, the median ridge of which persists, is seen in the 
late stage of the embryo. The premaxille are united on the palate; 
the premolars are large, displayed from within outward; the upper 
molars possess a well developed hypocone. The interdigital spaces are 
sharply defined proximally, and are without skin-folds on the ventral 
surfaces; the tragus is thickened on the inner border, and the external 
basal lobe lies within the conch. The sternum is keeled throughout. 
The ulna is provided with a shaft. The first metacarpal bone is flexed. 
These characters sharply separate Natalus from the Vespertilionide 
(Plecoti occasionally excepted). Some of the characters, such as the 
keeled sternum and fixed ulna, are found in other groups. 
The long bones bear a close resemblance to those of Lobostomidé 
(genera Mormops and Chilonycteris), and Macrotus among the Vampyri. 
The femur is quite similar, as seen in the flange on the inner side of the 
shaft near the head (best seen in Mormops) and in the backward posi- 
Proceedings National Museum, Vol. X V—No. 912. 
437 
