PROCEEDINGS OF THE UNITED STATES NATIONAL MUSEUM 
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SMITHSONIAN INSTITUTION 
U. S. NATIONAL MUSEUM 
Vol. 92 Washington: 1942 No. 3145 
NOTES ON BEETLES RELATED: TO PHYLLOPHAGA 
HARRIS, WITH DESCRIPTIONS OF NEW GENERA AND 
SUBGENERA 
By Lawrence W. Sartor 
Tue discussions presented and the conclusions reached in the present 
paper are the result of a critical study of 543 species of phyllophagan 
beetles from North, South, and Central America and the West Indies. 
These species are divided among four genera, with subgenera, as 
follows: (A) Phyllophaga Harris, 453 species (distributed in sub- 
genus Phyllophaga sensu stricto, 326 species; subgenus Listrochelus 
Blanchard, 56 species; subgenus PAytalus Erichson, 51 species; sub- 
genus Chlaenobia Blanchard, 12 species; and remaining subgenera, 8 
species) ; (B) Cnemarachis, new genus, 85 species (distributed in sub- 
genus Cnemarachis sensu stricto, 84 species, and Abcrana, new sub- 
genus, 1 species) ; (C) Clemora, new genus, 2 species; (D) Triodonyz, 
new genus, 3 species. 
About half of the species of Cnemarachis were studied at the United 
States National Museum, while nearly all the remaining 500 species 
are represented in my collection. 
In addition to the species enumerated above, I have studied well 
over a hundred species of Ancylonycha Dejean (synonyms: Holo- 
trichia Hope and Brahmina Blanchard), most of them undetermined, 
and also a large number of species of closely related genera (Hepta- 
phylla, Microtrichia, Haplidia, Encya, Empecta, Trematodes, Meta- 
bolus, Hoplochelus, Rhizotrogus, Lepidiota, and others) , some of which 
are valid and others apparently inseparable from Phyllophaga. 
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