BEETLES RELATED TO PHYLLOPHAGA—SAYLOR 161 
KEY TO THE NORTH AND CENTRAL AMERICAN 
SUBGENERA OF PHYLLOPHAGA 
1. Claws long and simple except that each has a distinct median, submedian, 
or basal tooth ; vertex never with a transverse carina; males of some species 
either with one spur connate with tibia or with only a single spur________ 2 
Claws either cleft (usually rather narrowly so) or serrate or pectinate or 
without teeth; vertex with or without a transverse carina; males with 
tibial spurs tree andlalways two dn number. 2) eee eee 4 
2. Dorsal surface with dense scales and with or without intermixed hairs. 
(Female with vestigial wings; male winged, its abdomen with compressed 
tubercles on each segment.) United States______ Tostegoptera Blanchard 
Dorsal surface without scales, vestiture consisting of hairs or lacking 
@MUINe Veen See a be ee et ee ee Seg 3 
3. Female with vestigial wings, males with functional (farcta) or not functional 
wings (cribrosa) ; metasternum of female very short, only as long as second 
abdominal segment; both tibial spurs free in male. United States and 
IY (ED: LY SS Aen ES ya eR ag ONY Serco a Eugastra LeConte 
Both sexes fully winged; metasternum in both sexes much longer than second 
abdominal segment; either both tibial spurs free in male or one connate with 
tibia or rarely one lacking. North, South, and Central America 
Phyllophaga Harris 
4. Claws serrate or pectinate, rarely quite simple; vertex usually with a strong 
to moderately strong transverse carina, rarely without a carina. , North and 
Aerts eATN CT Ct aa De Listrochelus Blanchard ° 
Claws never serrate or pectinate or simple, always variously cleft; vertex 
TONER UTED CELE) eh a y= Vamp eeencun ee a Pk a VL Tt ee Oy SE 5 
5. Male with hind claws widely cleft, claws of fore and middle tarsi simple. 
(Hemalecunknoyw7.)- . Mexico: sss ee ee ee Chirodines Bates 
Claws of all tarsi similar or nearly so in both sexes and narrowly to 
PA COU) Rial Ke) ES 5 eet pe a RU at Oe Ee Wee eee eee reer 6 
6. Tarsal segments densely to moderately pilose beneath, less densely so in 
females (segments broadened in males of several species) ; color usually 
light testaceous. North and Central America__--__ Chlaenobia Blanchard * 
Tarsal segments sparsely pilose beneath or not pilose (segments never 
widened in either sex); color variable. North, South, and Central 
PANTING Tol Gea ee ea Ba laa te ae an gee cece dae Phytalus Erichson ° 
GENOTYPES OF THE AMERICAN PHYLLOPHAGAN GENERA AND 
SUBGENERA 
Since the genotype designations of the American genera and sub- 
genera mentioned above are rather scattered, and also since in several 
cases designations have not heretofore been made, the following 
summary is presented : 
Phyllophaga Harris, 1826, Massachusetts Agr. Repos., vol. 10, p. 6. Type: 
Melolontha hirticula Knoch, designated by Glasgow in 1916. Present 
status: Accepted as a valid genus. 
@ See Saylor, Proc. U. S. Nat. Mus., vol. 89, pp. 59-130, 1940. 
7 See Chapin, Smithsonian Mise. Coll., vol. 94, No. 9, 20 pp. 1935, 
3 See Saylor, Proc. U. S. Nat. Mus., vol. 86, pp. 157-167, 1939. 
