BEETLES RELATED TO PHYLLOPHAGA—SAYLOR 165 
(Blanchard’s genotype designation in 1845 of Holotrichia pubera 
Dejean is not valid because “pubera” is a nomen nudum.) 
Holotrichia, which is in such general use today, falls unfortunately 
as an absolute synonym of Ancylonycha and is not available for use, 
since Hope designated as genotype I/. serrata Fabricius, the same 
species later designated as genotype of Ancylonycha by Duponchel. 
In our catalogs, Ancylonycha is listed as a synonym of Lachnos- 
terna Hope or Phyllophaga Harris and is credited to Blanchard, 
1845. In reality, the name Ancylonycha traces back to Dejean. In 
the second edition of his Catalogue, 1833, page 160, the first three 
species listed under this name are, in order: pubera Dejean (a nomen 
nudum), Melolontha leucophthalma Wiedeman, and Melolontha ser- 
rata Fabricius; listed in addition are 44 other “species.” In the 1836 
edition of the same Catalogue, on page 177, the first three names are 
given in the same order as in the 1833 edition, and in all 54 “species” 
are included, many of them nomina nuda. 
ANCYLONYCHA MINDANAONA Brenske 
Study of a series of reared adults of this species from Guam shows 
that the middle tibia is always without transverse carina and that 
the hind tibial carinae vary from nearly to entirely obsolete. In this 
character the species is close to the species of the West Indian genus 
Cnemarachis. The male genitalia are much more tubular than in most 
species of the continental American forms of the genus Phyllophaga, 
and the sexes of A. mindanaona are hardly separable because the sixth 
abdominal segment is not sexually modified. This last character 
alone, which it shares with the species of Rhizotrogus, separates it 
from the species of our continental Phyllophaga to which it is on the 
whole closely related. 
All data given above support the view elsewhere presented” that 
the genus Ancylonycha cannot be separated in toto from the continental 
American genus Phyllophaga on other than geographical grounds, 
though many of the species placed by various authors in the genus 
Holotrichia are at least subgenerically differentiated. However, the 
lack of sufficient material of determined species prevents a more 
conclusive statement of their relationship at present. 
1 Saylor, Rev. Ent., vol. 7, pp. 318-322, 1937. 
U.%. GOVERNMENT PRINTING OFFICE: 1942 
