( 499 ) 



ai)(loraeii the first, segment is mostly closely appressed to the thorax ; in Macro- 

 f/lossiim the first tergitc is reduced to a very narrow strip. The fau-tail is fonnd in 

 both sexes of a number of genera, sometimes only in the t? {I'Jiiri/pteri/x) ; it is a 

 Sesiad character indicated occasionally only by three small tufts {Nephele S S). 

 The scent-organ of the anterior coxa is sometimes strongly developed (Chromis). 

 Tlic mid- and hiudcoxa! merum are in the greater number of genera simple, or the 

 midcoxal merum is simjily carinate or subangulate ; but the large sharp tooth found 

 in Scxid and allies apjiears also in tliis tribe (Macrofflossunt and allies), though the 

 liiudcoxal merum is never as strongly jjroduced as in St'siu. 



The tibiae become spinose in several genera ; the apical thorn of the foretibia 

 fonnd in some ('t'phoiioiles (Sesiinae) and in many Spltii)gi<l(n', aseiminophorae 

 appears also here, with or without an additional armature of spines {OdontoHida ; 

 Proserjjiuus ; Arctonotus ; etc.). The midtarsal comb is very often lost respect- 

 ively reduced. The spurs may be long or short, unequal or equal in length : in the 

 latter case they are reduced. The proximal pair of the hindtiljia is lost only in 

 Microsphinr. The pulvillus and paronychium are rarely lost. The mid- and hind- 

 larsns of Macrofjlossum and allies is specialised in being compressed and having 

 ac(piired a dense spination on the outer surface. 



The ancestral antenna was doubtless spiuiform, long, and had an elongate 

 rongh-scaled end -segment. The clubbed antenna is a later acquisition. The short 

 end-segment found in numerous genera is a reduced long one, ;'.('. is a less 

 ireneralised character than the long segment. The length of the latter may become 

 exaggerated, as is the case in Acosneripc. The strongly clubbed antenna has a long 

 or a short end-segment. The bristles of the end-segment are sometimes much 

 prolonged, and resemble those of the Clioerocamjnnae {Fanacra; see also P/iiloi/ila) 



The pupa with compressed tongue-case (Choerocampid tongue-case) is a 

 derivation frnm an anteriorly cylindrical and obtuse pupa as found in Phlhtmpelicae. 

 On the other Jiaud, the Choerocamj>id pupa of Nephi'licae may become reduced, 

 assuming the aspect of the pupa of Sesiiiiue by losing the compressed projecting 

 tongue-case. Suidi Sesiad jiupae a]>pear frequently in genera with reduced head and 

 tongue of the imago {Deit/amia, J hirapsa, etc.). 



The pedigree inserted below gives expression to our view of the connection 

 between the various genera. There are three main branches : one leading from 

 Ikih'ph'da to hiirapsa ; the second from Neplielii to Rhopalopsi/cke ; the third from 

 Ni'pl/di' to Di'idnmia. We tiiought at first that this last branch miglit be a 

 development from tiie first. Rut on consideration of all characters we think to be 

 correct ia treating the third branch (on the left side in the pedigree) as a derivation 

 front some such form as Seplielc instead of Deilcphila. The qualifying remark 

 apjilied to the former pedigrees holds good also here. 



Key to the genera : 



(I. Spines of first row of abdominal 



tergites not longer than broad ... . I>. 

 Sjiines of first row of aljdominal 



tergites longer than bmad . . . . e. 



Ii. i'hid-segnient of uiitenna elongate, 



(^-antenna with fasciculated cilia(! <'1>. Mdcrotiloasiini. 

 Kntl-Hegnicnt of antenna elongate, 

 ci-aiiteiina sinjilar to those (jf V, 

 witiioni prolonged ciliae . CLI. Illiopiilojisip-lu'. 



