644 PROCEEDINGS OF THE NATIONAL MUSEUM. vol.40. 



testicles are numerous. The eggs become more or less separated 

 from one another by ingrowths from the wall of the uterus. 1 A para- 

 uterine organ develops in front of the uterus and into this the eggs 

 presumably pass and become inclosed in a single egg capsule. 



At least one important difference from Chapmania is exhibited 

 by Sphyroncotsenia uncinata, namely, the arrangement of the hooks 

 on the rostellum in ten to twelve rows. Less important is the lack 

 of hooks on the suckers of S. uncinata, which, moreover, may be due 

 simply to their loss in the specimens examined. The lack of a dorsal 

 longitudinal excretory vessel may be only an apparent and not an 

 actual condition. The location of the female glands toward the 

 pore side of the medullary parenchyma though different from the 

 condition in the type species of Chapmania is paralleled by the slight 

 displacement in that direction in C. tapika. The eggs in Sphyron- 

 cotsenia uncinata are apparently without parenchymatous or connec- 

 tive tissue envelopes outside the second shell, but it is possible that 

 such envelopes are present, as in C. tauricollis, the condition of the 

 material examined being such that this point could not be definitely 

 determined. 



As already noted, with regard to the uterus and para-uterine organ 

 S phyroncotsenia uncinata and Chapmania are very similar, the resem- 

 blance of S. uncinata to C. tapika being closer than to C. tauricollis 

 if Fuhrmann is correct in his statement that in the latter species the 

 uterus breaks down and the eggs become enveloped by parenchymal 

 tissue before they enter the para-uterine organ. 



The similarities in the structure of S. uncinata and Chapmania 

 indicate a relationship between them, but as the former is widely 

 different from the latter in the arrangement of the rostellar hooks, 

 the arrangement of these hooks being given great weight under the 

 present system of classification in the separation of genera throughout 

 the entire superfamily Tasnioidea, the necessity of erecting a separate 

 genus for S. uncinata is clearly apparent. 



S phyroncotxnia uncinata is much like Porogynia lata (Fuhrmann, 

 1901), the type and only known species of Porogynia Railliet and 

 Henry, 1909, though this species is at present classed in a different 



1 In C. tapika the cavity of the uterus becomes more or less separated into chambers, which, however, 

 always remain in communication, and contain groups of two to six eggs (Clerc, 190G). According to Fuhr- 

 mann (190S) the uterus in the genus Chapmania breaks down and the eggs become surrounded by paren- 

 chymatous capsules, after which they are pressed into a para-uterine organ which develops in front of the 

 uterus, and in gravid segments, after their release from the strobila, this para-uterine organ transforms into 

 a single large egg capsule. It would seem, however, both from Clerc's (1900) description and Fuhrmann's 

 (1909) description of C. tapika that in this species the eggs pass directly from the uterus into the para-uterine 

 organ. It may perhaps be considered an open question whether in C. tauricollis the uterus actually breaks 

 down, the eggs then becoming surrounded by envelopes of parenchymatous origin, or whether the uterus 

 as a result of extensive proliferation from its wall becomes more or less divided up into little chambers, the 

 outer envelopes of the eggs then being added as a secretion from or a direct transformation of the ingrowths 

 of the wall of the uterus. If the latter supposition be true, the eggs might apparently be contained in individ- 

 ual capsules, yet all of them still be within the cavity of the uterus, and completely surrounded by its 

 peripheral wall. 



