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The lateralis fibers destined to the raini buccalis and oticus enter the trigeminus ganglion 

 as a Single bündle, and while traversing the ganglion separate into two bundles, one destined to each 

 of the two nerves. The fibers that go to the ramus ophthalmicus lateralis do not traverse the ganglion, 

 lying, in sections, either whoUy separate and immediately dorsal to the ganglion or partly embedded 

 in its dorsal surface. This ophthalmicus bündle of lateralis fibers always traverses the skull either 

 through a partly separate part of the trigeminus foramen or through a whoUy separate but closely 

 adjacent foramen. 



From the trigeminus ganglion, in Scorpaena, the ramus ophthalmicus trigemini, the truncus 

 maxillo-mandibularis trigemini, the ramus communicans ad truncus hyoideo-mandibularis facialis, 

 and two independent branches arise; all of these branches issuing from the trigemino-facialis chamber 

 by its trigeminus opening excepting only the ramus communicans ad nervus facialis, which latter 

 nerve issues through the facialis opening of the chamber. Two or three bundles of communis fibers 

 traverse the ganglionic mass, one or two bundles going to the truncus maxillo-mandibularis and the 

 other one going toward the ramus ophthalmicus, but, so far as could be determined in my quite un- 

 satisfactory sections, going wholly to a branch that accompanies the ramus oticus lateralis. 



The ramus ophthalmicus trigemini arises from the anterior end of the trigeminus ganglion 

 by two Strands in all the sections of Scorpaena, Cottus and Lepidotrigla, one of these Strands running 

 forward dorsal and the other ventral to the ophthalmicus laterahs. From the dorsal one of the two 

 Strands, in Scorpaena and Lepidotrigla, a small branch, apparently a purely general cutaneous one, 

 is soon sent through the alisphenoid with the lateralis branch that goes to the small sixth organ of 

 the supraorbital canal, the two nerves being accompanied by a branch of the external carotid and 

 also by a branch of the vessel x. The general cutaneous component of this small nerve 

 would seem to be the homologue of one or both of the two meningeal nerves said by Herrick ('99, 

 p. 205) to have an extracranial origin in Menidia; and it may be added that no intracranial meningeal 

 nerves were evident in Scorpaena. Of these two nerves in Menidia, Herrick says, ,,I regard them as 

 primarily general cutaneous nerves" but ,,doubtless accompanied by sympathetic or other visceral 

 fibers." They are said to be ,, destined chiefly at least, for the skin of the top of the head", which 

 would seem to exclude them largely from the meningeal category. In embryos of Cottus this small 

 reentrant branch of Scorpaena and Lepidotrigla was not traced. 



After giving off this small reentrant branch, the two Ophthalmie nerves of Scorpaena and 

 Lepidotrigla, accompanied by the ophthalmicus lateralis, run forward dorsal to all the nerves and 

 muscles of the orbit, give off several branches while in the orbit, and then pass through the canal 

 between the frontal, mesethmoid and ectethmoid to reach the dorsal surface of the snout; their further 

 course not being investigated. In Ameiurus, Herrick ('Ol) says that the ramus ophthalmicus, his 

 supraorbital trunk, contains communis as well as general cutaneous fibers, which seems certainly 

 not true of the mail-cheeked fishes. According to Sagemehl ('84b, p. 71) the ramus ophthalmicus, 

 in the Characinidae, perforates the ectethmoid by a special canal; which is also not true of any of the 

 mail-cheeked fishes I have examined, nor of Scomber either. 



The truncus maxillo-mandibularis separates, as usual, into maxillary and mandibular portions, 

 the former being accompanied by the buccalis lateralis. A small branch is given off before the truncus 

 separates into its two parts, and running upward and backward innervates first the levator arcus 

 palatini and then the dilatator operculi. The further course and distribution of the nerve was not 

 investigated. 



