— 86 — 



of and does not traverse the trigeminus ganglion. Immediately after issuing from the skull it gives 

 off a small branch which turns upward and inward and, perforating the alisphenoid, enters the cranial 

 cavity. There it runs upward along the inner surface of the skull, lying in the membranous tissue 

 that lines that surface, and, piercing the frontal, reaches the small sixth organ of the supraorbital 

 canal, which it innervates. It is accompanied by a delicate branch of the trigeminus, apparently 

 of general cutaneous origin, as already stated, but this branch could not be aeparately traced in the 

 cranial cavity. In Lepidotrigla the same mixed nerve is found. In Cottus scorpius a similar nerve 

 is also found; but in the sections of this fish the nerve is lost beneath the terminal tube of the supra- 

 orbital canal, and no organ is evident in that tube. In the adult Cottus octodecimospinosus the 

 nerve is found innervating the terminal organ of the supraorbital canal, as in Scorpaena and Lepi- 

 dotrigla. Slightly posterior to this nerve, in Cottus, and having a closely parallel course, there is 

 a communis nerve which has an intracranial origin from the communis ganglion, and from there 

 runs upward and then backward along the side wall and roof of the cranial cavity. Whether or not 

 this nerve anastomoses with an intracranial branch of the vagus could not be determined in Cottus 

 scorpius, but in Cottus octodecimospinosus it does anastomose with such a branch and then issues 

 near the bind end of the skull; this seeming to indicate that it must form the anastomosis in Cottus 

 scorpius also. The nerve is accordingly the ramus lateralis trigemini of Stannius's descriptions, called 

 by Herrick, in his descriptions of Menidia, the facialis root of the ramus lateralis accessorius. It is 

 said by Stannius not to be found in Trigla, which I confirm. It is also not found in Scorpaena or 

 Dactylopterus. 



In the Cod, the facialis root of the lateralis accessorius is said by Herrick ('00) to be present 

 and to be accompanied by a few coarse fibers from the lateral line ganglion, which fibers would accor- 

 dingly seem to correspond to the branch to organ 6 supraorbital in Scorpaena. Whether or not this 

 latter branch of Scorpaena is represented in Pleuronectes by any part of Cole & Johnstone's ('Ol, 

 p. 128) ramus lateralis accessorius facialis, I can not decide. In Ameiurus it is represented in Herrick's 

 ('Ol) branch N. 5, which has a wholly intracranial course until it pierces the frontal to reach its organ; 

 a branch of the nerve going to the anterior head line of pit organs. 



The ophthalmicus lateralis, in Scorpaena, having given off the branch to organ 6 supraorbital, 

 runs forward between the two Strands of the ophthalmicus trigemini and soon gives oif a branch 

 which, running upward, separates into two parts and supplies the fourth and fifth organs of the supra- 

 orbital canal. This branch, in the several specimens of Scorpaena examined, and as already stated, 

 sometimes separated into its two parts before it pierced the frontal, sometimes so separated while in 

 its canal in the frontal, and, in one specimen, entered the supraorbital canal as a single nerve which 

 then, lying in the canal, supplied, first, organ No. 4 and then organ No. 5. After giving off this branch 

 the ophthalmicus lateralis continues forward, sends branches in succession to the third and second 

 supraorbital organs, and then supplies and terminates in the first organ of that line. 



The ramus oticus, in Scorpaena, separates from the buccalis while that nerve is traversing 

 the trigeminus ganglion, and issues from the ganglion as an independent lateralis branch but accom- 

 panied by two bundles of fibers one of which is of general cutaneous and the other of communis origin. 

 This latter bündle issues from the cranial cavity through the trigeminus foramen, and, so far as my 

 quite unsatisfactory sections show, contains all the communis fibers that go to the trigeminus excep- 

 ting those that accompany the truncus maxillo-mandibularis. These three bundles of fibers, united 

 to form a single nerve, run upward and laterally along the hind wall of the orbit and, traversing the 



