Poae| ORTMANN— AFFINITIES OF CAMBARUS. 123 
east, parallel to the strike of the mountains. This species, how- 
ever, has also been reported from Indian Territory (Ozark region). 
This locality is very strange, and at present is not connected with 
the main range, no localities being known in Missouri, Arkansas 
or the larger part of Tennessee (except the eastern extremity). 
But it is possible that a connection exists here, and if this should 
be so, this would indicate, as has been said above (p. 121) that the 
Ozark region is to be included in the original home of the sub- 
genus. C. monongalensis apparently is a representative form of C. 
carofinus in southwestern Pennsylvania. 
The most puzzling distribution is offered by the remaining three 
species, of which C. dogenes is the most widely distributed. This 
species has an eastern and a western range on both sides of the 
Allegheny Mountains. Apparently it has descended from the 
mountains, that is to say, represents a more highly specialized 
branch of the original mountain-loving chimney-builders. It has 
descended into the Atlantic coast plain on the one side, and is 
found from New Jersey to North Carolina (Cape Fear). On the 
other side, it has descended westward, and is found from south- 
western Pennsylvania over all the states north of the Ohio (also in 
Kentucky) as far north as Minnesota and Wisconsin, westward to 
Iowa (also reported from southwestern Wyoming and Colorado), 
Kansas, and southward to Louisiana. ‘This immense distribution 
represents possibly the widest known range of any of the species 
of crayfishes of the United States. The question remains open, 
whether the eastern and western range of C. dogenes is actually 
connected across the mountains. 
Of the other two species, C. wh/eri clearly is a local form of C. 
diogenes, inhabiting the sea coast (brackish and salt marshes) in 
Maryland. C. argzllicola is morphologically very closely allied to 
C. diogenes, and might be regarded, at least in Ohio, Michigan and 
Canada, asa local form developed at the northern edge of the range 
of C. dogenes. But the fact that C. argz//icola is also found in cen- 
tral and southern Indiana, in southern Illinois, and that it has been 
reported from Mississippi and southern Texas (Victoria and Bra- 
zoria), does not render this assumption probable: further investi- 
gations of the range of these two species (@ogenes and argz/licola) 
in the south and west are desirable, before their mutual geographic 
relation can be ascertained. 
PROC. AMER. PHILOS. SOC. XLIV. 180. I. PRINTED JULY 29, 1905. 
