[Vor. 1 
390 ANNALS OF THE MISSOURI BOTANICAL GARDEN 
in pl. 21 fig. 11. It will be seen that there is no indication of a 
rosette, and the branching is quite different from that of Œ. 
mut. rubrinervis. In many cases, however, a rosette is formed. 
When the rosette is omitted the branching is changed. Plate 
21 fig. 10 shows on a larger scale another individual in flower. 
The stem-leaves differ from those of GZ. mut. rubrinervis in being 
narrower, more pointed and smoother. 
In this race the red papille on the stem were very numerous, 
and the buds likewise were slightly more red than in (7. mut. . 
rubrinervis. The modal color pattern of the whole population 
was 6 as in Œ. mut. rubrinervis, but plants with their mode at 
7 were much more numerous than in the latter (see Gates, '11, 
p. 351). The race as à whole inherited the capacity for pro- 
ducing a slightly greater amount of pigment. The ovary usu- 
ally bore many long hairs arising from red papille; on the hy- 
panthium were few long hairs from slight green mounds; and 
on the bud cone scattered long hairs from conspicuous red 
papille. In occasional buds, when the color pattern was only 
8, the green papille were more numerous. Іп addition to the 
color pattern of the sepals there was usually weak red on the 
hypanthium.! 
The same conditions as regards pigmentation have been main- 
tained in later generations. Тһе plants were, however, by no 
means uniform in all respects, and this was not to be expected 
since they were derived from one individual of a freely inter- 
crossing population. Plate 21 fig. 9 represents a rosette of 
one of the Е, plants. The latter differs obviously from the one 
represented in pl. 21 fig. 8, but the race retained in this and 
subsequent generations the long, narrow, smoothish leaves as 
well as the pigmentation. The various Ез and Е, families, each 
derived from a selfed individual, produced sub-races differing 
more of less from each other and varying within narrower limits. 
It does not appear that the Mendelian theory of the sorting out 
of factors, or "genes," affords an adequate explanation of all 
these phenomena. 
1 Since this condition of bud-pigmentation resembles that obtained in certain F4 
and Ез hybrids of Œ. mut. rubricalyz and Œ. grandiflora (see Gates '14), it is possi- 
ble that it may have arisen in a similar way, i. e., by the appearance of a red-budded 
mutation which subsequently crossed with other species, in which crosses some blend- 
ing of pigmentation occurred giving rise to the present condition. 
