1921] 
PAYSON—MONOGRAPH OF THE GENUS LESQUERELLA 107 
seems no other alternative than to consider this broadened con- 
dition of the filament as another character possessed by the 
ancestral type of the genus. 
The Nectar Glands.— European botanists have for many years 
been discussing the taxonomic merits of a study of the nectar 
glands in the Cruciferae, with quite diverse resulting opinions. 
Much yet remains to be done before their value may be estimated 
satisfactorily, but it is certain that they must not be neglected 
in any work, particularly in groups above generic rank, that 
seeks to unravel natural relationships. Because of the small 
size and lack of distinguishing color, the diffieulty attendant 
upon their investigation prevents their use in diagnoses or keys 
intended for popular use and greatly hinders their investigation 
by the specialist. The degree to which they are developed 
varies considerably between species and also ap- 
parently between individuals, but within the 
genus the general plan of their arrangement re- 
mains fairly constant. These glands consist of &@} 
elevations of secretive tissue on the receptacle Í 
near the bases of the stamens. In Lesquerella 
the greatest development occurs in the immedi- 
ate vicinity of the base of the solitary stamen ig. Diagram 
and from here extensions reach toward the bases — -— ears 
of thedouble stamens, as shown in the diagram- -- an in L. au- 
matic sketch of the location of the glands in L. 
auriculata (fig. 1). Occasionally there is a considerable growth of 
glandular tissue below or between the double stamens. Some- 
times the ring surrounding the solitary stamen may be unbroken 
on either side or may be open on the opposite side from that 
shown in the diagram. The glands may be rounded elevations, 
ridge-like or plate-like masses in different species, or they may 
be even produced into short, horn-like processes as in L. lasio- 
carpa. In general, the greatest glandular development is to be 
found in the more primitive species, and probably the glands of 
the group from which Lesquerella was derived were not unlike 
those of L. auriculata. 
The Pedicels.—Considerable importance attaches to the form 
of the pedicels in the determination of species because of three 
characteristic and relatively constant positions assumed. The 
