1922] 



PAYSON — STUDY OF THELYPODIUM AND ITS IMMEDIATE ALLIES 299 



Gard. Herb.) ; sand bluffs of the sea-shore, San Diego County, 

 April 25, 1882, Pringle (Mo. Bot. Gard. Herb.); Rose Canyon, 

 La Jolla, March G, 1914, Clements & Clements 30 (Mo. Bot. 

 Gard. Herb.); San Diego, 18S2, Parry (Mo. Bot. Gard. Herb.); 

 Encanto, 1882, Parry (Mo. Bot. Gard. Herb.) ; San Diego, April 

 25, 1903, Brandegee 3390 (Mo. Bot. Gard. Herb.) ; lower hills, 

 San Bernardino Mountains, March and April, 1888, S. B. & W. 

 F. Parish (Mo. Bot. Gard. Herb.). 



Although carried for many years as a species of Streptanthus 

 this plant has frequently been confused with other plants now 

 considered members of the genus Caulanthus. C. Coulteri, for 

 instance, was first segregated from material identified at one time 

 by Dr. Gray as S. heter vphyllus . In one place (Proc. Am. Acad. 

 6: 186) he remarks on the difference in the stigmas of these two 

 plants but allows that much intraspecific variation. In the 'Flora 

 Franciscana' Dr. Greene associates this plant with Streptanthus 

 Coulteri and S. Parryi (C. Lemmonii). Both of those plants 

 have been retained in Caulanthus for many years. The present 

 plant, C. heter vphyllus, has previously been retained in Streptan- 

 thus for two main reasons: it has pods that are somewhat com- 

 pressed and seeds that are narrowly margined. This is taken to 

 mean that C. heter vphyllus shows a parallel development to what 

 occurs in Streptanthus. From a phylogenetic standpoint it is out 

 of place in Streptanthus, since it is undoubtedly derived from ex- 

 isting species of Caulanthus — or a species very similar to existing 

 ones — and not from the same point that it is believed Streptan- 

 thus has developed. 



14. C calif ornicus (Wats.) Payson, n. comb. 



Stanjordia calif ornica Wats. Geol. Survey Calif. Bot. 2: 479. 

 1880; Prantl in Engler & Prantl, Nat. Pflanzenfam. III. Abt. 

 2: 206. 1891; Robinson in Gray, Syn. Fl. N. Am. I 1 : 171. 1895; 

 Hayek, Beih. Bot. Centralbl. 27 1 : 314. 1911. 



Streptanthus californicus Greene, Fl. Franciscana, 256. 1891. 



Annual, glabrous or very sparingly pilose near the base: stem 

 erect, branched from near the base and upwards, 2-4 dm. high: 

 radical leaves oblanceolate, sinuately lobed to pinnatifid, obtuse, 

 3-6 cm. long; cauline leaves ovate to oblong, amplexicaul, shal- 

 lowly dentate, obtuse: sepals purple-tipped, unequal, membran- 

 ous and saccate near the base, 7-9 mm. long; petals whitish, 

 little longer than the sepals, claw dilated, blade small, crisped; 

 filaments 5-9 mm. long, longer pair sometimes slightly united 



