Genera Cornufcr, Platymantis, <&c. 373 



of the latter, which agree with those of Discodeles (large 

 nasals in contact with the frontoparietals, omosternal style 

 forked at the base) and differ from those oi Amolops and 

 Staurois (small nasals widely separated from each other and 

 from the frontoparietals, omosternal style not forked). 



Digital discs absolutely similar to those of Cornvfer are 

 found in Staurois, Cope (type: Ixalus natator, Gthr.), and 

 in a frog from Kina Balu, North Borneo, described by me 

 thirty years ago under the name of Ixalus latopalmatus, 

 which I now regard as the type of anew genus, Simomantis, 

 so named in allusion to the curious pug-like form ot the 

 snout ; this frog agrees with the species grouped under Staurois 

 in the' very large digital discs, broader than long, supported 

 by T-shaped phalanges in which the horizontal limb is longer 

 than the longitudinal, in the absence of an intercalated bone 

 between the penultimate and distal phalanges, in the outer 

 metatarsals separated to the base by the very broad web ot 

 the toes, in the absence of vomerine teeth, and in the osteo- 

 logical 'characters referred to above. In addition to the 

 structure of the digital discs, Simomantis is distinguished 

 from Rana, as it is also from Staurois, by the webbed fingers, 

 a character which had led me to refer S. latopalmata to the 

 vicinity of Ixalus bimaculatus, Peters, likewise from Borneo, 

 which is, however, a true Ixalus. 



Simomantis is confined to Borneo ; Cornvfer is known 

 from Burma (C. tenasserimensis, Stoliczka, originally de- 

 scribed as a Ratio), Borneo (C.baluensis, Blgr. = Rana sariba, 

 Shelford), the Philippines (C.guentheri, B\gr., jagorii, Peters, 

 worcesteri, Stejneg., corrugatus, A. Dum.), New Guinea 

 (C. unicolor, Tscliudi), the Solomon Islands (C. guppyi, 

 Blgr.), and the Fiji Islands (C. dorsal/*, A. Dum., inter- 

 medins V. Mull.) ; whilst Plntymantis is represented in the 

 Philippines {P. corrugata, A. Dum., meyeri, Gthr.), Xew 

 Guinea and neighbouiing islands [P. corrugata, A. Dum., 

 punctata, Peters & Doria, beauforti, v. Kamp.), Now Britain 

 (P. boidengeri, Boettg.), the Solomon Islands (P. solumonis, 

 Blgr.) and the Fiji Islands (P. vitiana, A. Dum., uni/ineata, 



Peters). 



Although the presence or absence of vomerine teeth is 

 o-enerally^regarded as of generic importance, it would be so 

 obvious a violation of the principles of natural classification 

 to separate genetically Staurois hainanensis, Blgr., from 

 Rana latopalmata, Blgr. (Amolops afghana, Cope) , or Staurois 

 natator, Gthr., from Rana guttata, Gthr., on this ground, that 

 1 have no hesitation in referring Slaxirois hainanensis to Rana, 

 in spite of the absence of vomerine teeth, and 411 modifying 

 the definition of the genus Staurois, founded on the absence 



