382 Mr. R. I. Pocock on some 



differences between the hind feet of Danis and Ursus may 

 be seen by comparing my sketch of the right foot of the 

 former with that of the latter, published by Boas in 1909 

 (Zool. Atiz. xxxiv. p. 529). This figure shows the digits of 

 U. arctos in their natural position and not separated to 

 their full extent as in my figure of the feet of I), horrid} Its. 

 Also in Boas's figure it may be noted that the pad of the 

 first digit is set farther back, a much greater extent of the 

 heel is covered with hair, and that the transverse groove on 

 the inner half of the sole is larger and invaded by hair. 

 This character, however, is variable in U. arctos. 



A point of special interest connected with the feet of 

 I >tinis horribilis is their resemblance in the alignment and 

 fusion of the digital pads to the feet of Mel ursus ursinus. 

 In the latter, however, all the digital pads are united to the 

 same, or nearly the same, extent * as are those of the third 

 and fourth digits of the hind foot in D. horribilis, and the 

 posterior border of the pads is less well defined and the area 

 between them and the plantar pad is quite naked. Similarly, 

 in the fore foot of Melursus the area between the plantar 

 pad and the carpal pad is naked, and above the carpal pad 

 the integument is for a short distance quite scantily clothed 

 with hair. Nevertheless, the structure of the feet of Mel- 

 ursus suggests that this genus is a specialized modification 

 of the Dunis-type rather than of the Arcticonus thibetanus or 

 Helarctos malay anus-type, the hemispherical ulnar carpal 

 pad and reduced radial carpal pad recalling these structures 

 in Danis, Ursus, and Euarctos. 



In Helarctos malayanus, as I nave already stated from 

 an examination of living specimens and dried skins, the feet 

 resemble tolerably closely those of Arct/conus thibetanus. 

 This I have been able to verify on a fresh specimen from 

 British North Borneo. The digital pads are free and 

 susceptible of wide divarication as in Euarctos, but when in 

 contact their alignment is not quite so strongly curved as in 

 that genus. The hair clothing the area between the digital 

 and plantar pads is reduced to lour patches opposite the 

 interdigital spaces, and these patches are much larger on 

 fore than on the hind feet and the anterior border of the 

 plantar pad is less well defined than in Euarctos. In the 

 lore foot the carpal pad forms an area as wide as the plantar 

 pad and continuous with it, as in Arcticonus thibetanus, but 

 the divisional line between the two is much less emphasised 

 than in the specimen of that species 1 figured in 1911. In 



* Tii of fusion varies within the genus, the sutural line 



sometimes being distinctly retained, sometimes almost ohliterated as in 

 my original figure. 



