Mr. R. I. Pocock on the Genus Eupleres, &c. 349 



pads is quite naked. The plantar pads are large and normally 

 trilobate, and the internal lateral lobe has a moderately large 

 pollical {pi.) or hallucal lobe {hi.) attached to its posterior 

 angle. There is no metatarsal pad, but a strip of naked skin 

 runs obliquely inwards and upwards to the middle line from 

 the angle of the external lateral lobe of the plantar pad. The 

 carpal pad {c.) is double and closer than usual to the plantar 

 pad, its subspherical external or ulnar moiety being larger 

 than the internal ; between the former and the external 

 lateral lobe of the plantar pad there is a small area of naked 

 skin. 



In the characters so far enumerated there is nothing to 

 distinguish the feet of Eupleres, otherwise than generically, 

 from those of the genera previously discussed in this paper, 

 or, indeed, from those described in my paper upon the 

 Viverrina? (P. Z. S. 1915, pp. 131-149) ; but in one character 

 they are peculiar, namely, the comparatively large size and 

 low position of the hallux and pollex — a primitive feature, 

 suggesting that, although truly digitigrade, Eupleres, when 

 standing, has the five digital pads instead of four in contact 

 with the ground. 



Conclusion. 



Assuming provisionally the absence of perineal scent- 

 glands in Poiana, the four genera discussed in this paper 

 differ by that negative character from those I have recently 

 dealt with elsewhere, which may be referred to the four 

 subfamilies of Viverridae, namely, the Viverrinaa (Viverricula, 

 Viverra, Civettictis, Genetta), the Hemigalinas (Hemigalus, 

 Ghrotogale, Diplogale), the Cynogalinse {Gynogale), and the 

 Paradoxurinae {Paradoxurus, Paguma, Macrogalidia *, Arc- 

 tictis, Arctogalidia, N andinia) . 1 do not think it is placing 

 a too high value upon the characters distinguishing these 

 groups to assign them the rank of subfamilies. The Para- 

 doxurinse, indeed, may be susceptible of finer subdivision, 

 Nandinia especially having strong claims to be regarded as 

 the representative of a special group of that standing. How- 

 ever that may be, the fourteen genera so classified agree, so 

 far as is known fj with each other in possessing the scent- 

 gland in diverse forms and positions in both sexes \ ; and 

 this is a very special organ probably inherited from a common 



* Proposed by Sdrwarz (Ann. & Mag. Nat. Hist. (8) v. p. 423, 1910) 

 for Paguma mussenbroeki from Celebes. 



t Not verified in Chrotogale and Diplogale. 

 i Not verified in the male of ArctogaHdvt. 



